Debbie McGee has said she and husband Paul Daniels lived a "fairytale life" together.

The magician died on Thursday aged 77 at his Berkshire home, a month after being diagnosed with a brain tumour.
McGee said her husband did not know he was dying in his final days.
"He was in, what the consultants said was, a happy, confused state. He knew who everybody was, but his brain wasn't processing new information [due to the tumour]," she told the BBC.
McGee, who married Daniels in 1988 after many years as his on-stage assistant, said they had the kind of marriage "most people dream of", saying: "We had a life that was full of laughter... we were so happy together."
Daniels was diagnosed with an inoperable brain tumour last month.
"I've cried my heart out some nights but it all happened so quickly," McGee said.
"On the day he was diagnosed the tumour had already spread so far that they knew he only had a few weeks to live. Then, in two days he deteriorated from feeling lethargic to having other symptoms, so he actually didn't really know what was going on.
Daniels was born Newton Edward Daniels in Middlesbrough and developed his magic skills in working men's clubs, making his TV debut on talent series Opportunity Knocks.
He then took over the primetime Saturday night slot with his BBC show, which started in 1979.
Daniels was known for a string of catchphrases, including the line: "You'll like this... not a lot, but you'll like it."
BBC correspondent Nick Higham said: "Paul Daniels could be chippy and some thought him smug. He fell out with BBC managers who dropped his show in the 1990s.
"But there was no doubting his popularity - or his talent."
It was initially suspected Daniels had suffered a stroke after he fell at home, but it was then discovered he had an inoperable tumour.
His son, Martin Daniels, said at the time: "He has said before, 'When it's your time it's your time' and that's how he is trying to face up to things."
As well as McGee, Daniels leaves his three sons from a previous marriage - Martin, Paul and Gary.
Gary Daniels has tweeted a picture of a rabbit in a magician's hat with a tear in its eye, by friend and artist Helen Martin, and wrote: "It is with incredible sadness that I can confirm that Dad, Paul Daniels, passed away overnight."
In addition to his magic, Daniels presented game shows in the 1980s and '90s, including Wipeout, Every Second Counts and Odd One Out.
He also designed special effects for the stage productions of Cats and Phantom Of The Opera, and was commissioned by the English National Ballet to create magical effects for The Nutcracker.
Daniels had continued touring with his magic show and was booked to perform at venues across the UK this year, according to his website, before news of his illness emerged.
He supported a number of charities during his life, taking part in fundraising events and entertaining children at Great Ormond Street Hospital.
The Magic Circle said Daniels had "been an inspiration to many magicians" and is of "legendary status" in the world of magic.
In a statement, it said: "At the height of his career The Paul Daniels Magic Show on BBC One pulled more than 20 million viewers - that was more than one third of the British population regularly tuning in on a Saturday night to watch a magician  make them laugh and witness feats of wonder such as his famous Chop Cup routine.
"In this day and age of fragmented media platforms, no other magician is likely to be able to achieve such a TV ratings record."
It added that "the magic world owes Paul a huge debt".
Illusionist Dynamo told the BBC: "Without Paul Daniels I don't think I'd be doing magic today, so it's really a testament to how important he is for the magic world, he was a true master and [his death] is just such sad news.
"He was a real personality, he didn't just perform a bunch of tricks, he knew how to take something and make it magical but humorous at the same time.
"He had his own style which I don't think anybody could ever emulate or recreate. Paul Daniels for me is probably the best magicians Britain has ever had. He has such an amazing legacy in magic."
Louis Theroux, who filmed a documentary with Daniels and McGee, wrote: "RIP Paul Daniels, a thoughtful skeptic, enemy of hucksterism and paranormal flimflam, and gifted magician."
Ola Jordan, who danced with Daniels when he appeared on Strictly Come Dancing, tweeted that it had been "a pleasure and honour" to work with her "amazing dance partner".


Debate will continue about whether this was the UK's first 'social media election' but party leaders, candidates and even the electorate have certainly been online  to campaign, comment or simply make a joke.

One recurring social media prop has been the use of images that aren't quite what they seem.
Key moments in the campaign have been immortalised with some very creative photoshopping, which frequently led to the birth of widely shared memes and hashtags and generally resulted in carefully staged campaign 'moments' not having the entirely desired effect...
The campaign kicked of with a slightly unusual celebrity endorsement of the Lib Dems by reality TV star Joey Essex, who told wide-eyed journalists he had thought the party was called the Liberal 'Democats' before meeting leader Nick Clegg who helpfully put him straight.  The Lib Dems took it in good humour and took to photoshop themselves to join in the fun, briefly changing their website logo...
When Ed Miliband appeared next to a huge stone slab with his election promises engraved on it, it was open season for photoshoppers everywhere as social media lit up with Moses jokes and it quickly became known online as #EdStone.
Shadow Chancellor Ed Balls was roundly mocked online as the fourth anniversary of #EdBallsDay, the day that the internet marks his accidental tweeting of his own name, rolled around and he was photoshopped into all manner of inappropriate places.
Ed Miliband became an unlikely online pin up as the his face was photoshopped onto images of famous movie stars, musicians and footballers bodies and #milifandom was born, some taking it more seriously than others.
One by one as the major parties manifestos were launched they were mercilessly mocked online.
And then there was the dead pigeon incident... where an unfortunate pigeon fell victim to the Lib Dem campaign bus and Twitter reacted, well, as you might expect it to...
The Tories weren't safe from photoshop fiends either, as they (and all the parties) tried to convince us that they are inclusive and forward-thinking.  We looked at  the kind of political rebrands social media generously offered the main parties, the Tory tree getting more than one reworking.
As the campaign rumbled on David Cameron's newfound passion didn't go unnoticed as he started to appear at events mid-campaign with his sleeves rolled up, and for once it was the audio rather an the imagery that got a reworking. Remarking at one event that the campaign was making him feel 'pumped up', this clip from his speech which was quickly turned into a mash up with the Black-Eyed Peas song of a similar name.
There were however some stories that made headlines where no photoshopping or re-mastering was required...
The Green Party offered an alternative vision with an advert portraying the Conservative, Labour, Lib Dem and UKIP leaders as a slightly questionable boyband. On the other hand if teenage girls got onboard with #milifandom who knows what will win votes.
Staying on the musical theme #constituencysongs was an early Twitter hit, when tweeters took to the site to offer song titles reworked to contain constituencies, to the amusement of bored journalists everywhere.
Again no tweaking required, when an enthusiastic UKIP voter in Essex when asked his name by a journalist replied:
As a frightening number of error ridden leaflets appeared on people's doormats, they took to social media to share their horror, altering the images little further than an indignant biro-ringing of the offending error.
Yes, that's a stick with a wig on it. No, it hasn't been photoshopped. It's one of a number of unusual images that we were treated to, via social media, as we took a look at the weird world of hustings.
The unusual paring of Ed Miliband and Russell Brand appeared on our screens leading to... you guessed it, #milibrand as Ed appeared to 'rebrand' himself, adopting an estuary twang for the duration of the encounter.
You might not believe it but these pictures genuinely weren't photoshopped. Yep, they really were all wearing the same thing, while telling us earnestly how different they all were from the other guy.  But as ever the truth will out (on social media).
And wait for it... The UKIP ringtone. You didn't think we'd forgotten that gem did you? No re-mastering required this treat is 100% original. You're welcome!
Written by Kerry Alexandra


Australian Nick Kyrgios has told a group of opposing fans to "man up" after swearing at them in his win over Israeli Dudi Sela in Acapulco.

Kyrgios, 21, reacted after apparently being disturbed as he prepared to serve in Tuesday's 3-6 6-3 6-3 win.
The world number 17 later laughed off Australian broadcaster 7 News' report that four fans want an apology.
Sela confronted Kyrgios after the match at the Abierto Mexicano Telcel, with the umpire coming between the players.
This content will not work on your device, please check Javascript and cookies are enabled or update your browser
The incident is the latest in a string of controversies for Kyrgios who accused an umpire of being biased in his semi-final defeat by Jo-Wilfried Tsonga in Marseille last week.
He was banned for eight weeks and fined for his on-court behaviour at October's Shanghai Masters, although he accepted a 'care plan' agreement that would reduce the suspension to three weeks.
Kyrgios will play former world number one Novak Djokovic in the last eight from 04:00 GMT on Friday morning.
Media playback is not supported on this device


Worcester Warriors have signed winger Tom Howe from fellow Premiership side Wasps for the 2017-18 season.

The Buckinghamshire-born youngster, 21, has been associated with Wasps since their High Wycombe days, when he joined from Beaconsfield.
Howe, who has two tries in 13 first-team appearances, played for England's Under-20s as they won the Junior World Championship title in 2016.
"Tom is a hungry, young player," said Warriors director of rugby Gary Gold.
"He will add further depth to our options in the back three. He has shone for Wasps when presented with the opportunity. Hopefully, he can use the resources around him at Sixways to flourish."
Howe has also had loan spells in the Championship with Jersey Reds, and in National League One this season at Coventry.
Wasps have also allowed a player to join Warriors' Premiership relegation rivals Bristol.
Loosehead prop Tom Bristow has joined Bristol for the rest of the season, making him available for Sunday's Worcester-Bristol match at Sixways.
Worcester will start Sunday's game two points ahead of Bristol, who beat them when the two sides first met this season at Ashton Gate on Boxing Day.
Meanwhile, Wasps back-row forward Alex Rieder has signed a new, undisclosed-length contract with the Premiership leaders.


When the sorry story concluded under a pink Adelaide sky, it was hard to remember that it has not always been this way.

The clatter of James Anderson's stumps and the blur of green Bangladesh celebration marked the end of another disappointing World Cup campaign, but there was a time when England were quite good at one-day cricket.
In the tournament's very first match, at Lord's in 1975, England got so many runs that India didn't even try to chase them. Three final appearances would follow in 1979, 1987, and 1992.
The 1992 tournament, held in Australia and New Zealand, was the competition that invented modern one-day cricket - coloured clothing, white balls, pinch-hitters, innovation. It was also when England's success ended.
Since then, they been eliminated in the first round of the World Cup on three occasions and have failed to win a single match in the knock-out stages.
With yet another period of soul searching and "blank sheet of paper" planning under way, how can England fix their one-day team in time for a home World Cup in 2019?
Despite their calamitous exit, England have in batsmen Jos Buttler, Joe Root, James Taylor and Moeen Ali a handful of players on which a one-day team could be built.
But how do they replace opening bowlers Anderson and Stuart Broad, both of whom seem unlikely to be playing ODIs in 2019?
All-rounder Ben Stokes is certain to come back into the reckoning, maybe with Kent batsman Sam Billings and Yorkshire leg-spinner Adil Rashid, but there is a paucity of fast-bowling options in the county game.
"We have good players and we need to give them time to develop," said former England captain Alec Stewart. "Stokes, Billings and [Surrey batsman] Jason Roy are game changers. They will make errors but they will develop.
"Can we get those three into the England set-up and give them the freedom to do what they do for their counties?"
England played nothing but one-day cricket for five months in the run-up to the World Cup, even altering the Ashes schedule to aid preparations by touring Australia last winter.
That still did not stop them from making more major changes for the opening games against Australia.
Taylor, who had enjoyed success at number three, was dropped down to six to accommodate Gary Ballance, a man who had not played a one-day international since September. Broad opened the bowling instead of regular opening bowler Chris Woakes.
"It wasn't a World Cup that sneaked up on us and said 'boo'," said former England pace bowler Matthew Hoggard.
"We shot ourselves in the foot by changing the captain - Alastair Cook was replaced by Eoin Morgan - a short period before the World Cup and we didn't have a settled battling line-up.
"We have to look at the structure of our 50-over cricket and that has to stem from the coach and the way we want to play it."
The record-breaking run scoring of the 2015 World Cup demonstrates that 50-over cricket is emulating Twenty20 cricket in more ways than white balls and coloured clothing.
Yet few members of the England's one-day team take part in domestic T20 cricket, either at home or in high-profile tournaments like the Indian Premier League or Australia's Big Bash.
Since the beginning of 2013, Anderson has played four domestic T20 games anywhere in the world, Broad three, Ian Bell two and Root just one.
Compare those to the numbers of New Zealand's Brendon McCullum (27), India's Virat Kohli (30), South African duo AB de Villiers (34) and Dale Steyn (39) and Australia's Mitchell Johnson (34),
In a cricketing world of T20 globetrotters, only five of England's 15-man party have played in a T20 league outside the UK.
Some will point to the crowded nature of England's Test calendar, but players from other countries with similar schedules find time.
In that same period since January 2013, Broad, Bell and Anderson have played 22 Tests, Root 21. McCullum has played 22, Kohli 19, Steyn and De Villiers 18.
"We have to give our best players the chance to play in the right environment, to play in the IPL and Big Bash without restrictions," said former England fast bowler Steve Harmison.
"Unless we do that, the rest of the world will get better as we will stand still."
Making time for English players to take part in the Indian Premier League is fine in theory, but they have to be wanted. Nottinghamshire batsman Alex Hales, for example, went unsold at the latest IPL auction.
If that is uncontrollable, at least the England and Wales Cricket Board can aim for the highest possible standard of limited-overs cricket in the UK.
Counties only returned to playing 50-over matches in line with ODI playing conditions in 2014, having previously dieted on 40-over contests.
In addition, a protracted T20 season, taking place over the whole summer, means it is harder to attract the best international players to take part.
There have been frequent calls from the likes of Michael Vaughan and Kevin Pietersen to introduce a T20 franchise system similar to the IPL or Australian Big Bash.
"We need a more intense one-day competition," said former England batsman Geoffrey Boycott. "Australia and South Africa have a more fierce system.
"Ours seems low key, it doesn't mean quite as much. If you play in a more intense, competitive atmosphere, it stretches you and makes you a better player."
Moores has had success in the rebuilding of the England Test side. Of the XI that came from behind to earn a 3-1 series win against India last summer, six had not won a Test match before, let alone a series.
Moores's predecessor Andy Flower decided that the Test and limited-overs was too much for one man, and split the duties with Ashley Giles.
Giles had mixed results, taking England to the 2013 Champions Trophy final, but losing to the Netherlands as part of an early exit from the 2014 World T20 before leaving his role in April 2014.
If the job was to be split again, might England be able to attract a high-profile international name who could combine the role with a lucrative contract in the IPL or Big Bash?
New Zealand's Stephen Fleming and South African Gary Kirsten have been mentioned, as has Yorkshire's Australian coach Jason Gillespie.
"They need some foreign input," said BBC cricket correspondent Jonathan Agnew. "If it's only for limited-overs cricket, then they might have a chance of getting a big name like Stephen Fleming or Gary Kirsten.
"They need someone who understands English cricket, but also has coached elsewhere in the world and knows how modern one-day cricket is played."
However, Giles says a "foreign coach isn't necessarily the answer".
"Stephen Fleming is a very good man and he's done well in the IPL, but we need to look from top to bottom how we play our cricket," he said.
Moores has talked a lot about "analysing the game data".
Such an approach adds weight to claims that England rely too much on statistics, a charge denied by Moores, Morgan and ECB managing director Paul Downton.
If we are to give them the benefit of the doubt, then the line has to be drawn at some of the indecipherable phrases trotted out throughout the tournament.
Media playback is not supported on this device
Morgan, for example, said this before the Bangladesh game: "Moving forward, focusing on our strengths as a unit, individually, is very important."
Contrast that to the straight-talking of South Africa paceman Steyn: "If they need seven off the last over, I'll make sure they score three. If we need two wickets to win, I'll get those two wickets."
Former England wicketkeeper and 2005 Ashes winner Geraint Jones says that England are "obsessed by figures".
"They look like a team that plays to figures," he added. "Look at the teams who've set the World Cup alight.
"Brendon McCullum just goes out there and tries to wallop everyone everywhere."
Alec Stewart, Matthew Hoggard, Steve Harmison, Geoffrey Boycott, Jonathan Agnew, Ashley Giles and Geraint Jones were speaking on BBC Radio 5 live.
Listen to highlights from Test Match Special's and 5 live Sport's 2015 World Cup coverage.


The last RAF aircraft to be repaired by the Ministry of Defence (MoD) at St Athan has flown out, ending 75 years of maintenance at the site.

The remaining 200 workers at the base will be made redundant after the VC10 left at 15:50 GMT on Thursday.
Vale of Glamorgan MP Alun Cairns called for the skills of the workforce to be kept within the area.
The Welsh government said it was working to attract new companies, with interest in a new enterprise zone.
Mr Cairns said he was sorry to see the last aircraft leave the site, but the "expert knowledge base" needed to be retained locally.
"I'm in touch with the MoD regarding the plans for the super hangar and am optimistic about the outcome," said the MP.
Paul Lindsay of the Wales Aerospace Forum said it was "a sad day" and "a great disappointment" for the local community.
"There are positives in that we've had 75 good years," he said.
"But the disappointment is that between the Welsh government and the MoD they have invested £240m there over the past 10 years.
"Let's hope that collectively they decide what the future is for St Athan. The Welsh government and the MoD are working hard to fill the gap."
St Athan's history as an aircraft maintenance base dates back to 1938.
Just before and during World War II many different types of aircraft were assembled, test flown and maintained there by a staff of 14,000, and the base inevitably became a target for German bombers.
By the mid-1990s there were 3,500 engineers repairing fighter jets.
But since then the St Athan site has since endured more than a decade of uncertainty and failed attempts to secure skilled jobs.
In 2000 the Red Dragon Project began, aimed at modernising the ageing MoD facilities there, along with the construction of a £77m super hangar for fast jet repair.
The main tenant for the new super hangar was to be the Defence Aviation Repair Agency, Dara.
But in 2005, UK ministers announced they were closing the fast jet business at St Athan, with the loss of hundreds of jobs.
In March 2009 the National Audit Office and the Wales Audit Office criticised both the MoD and Welsh authorities after it had cost the taxpayer £113m and failed to deliver thousands of jobs.
The then Defence Minister Quentin Davies revealed in November 2009 that nearly 340 jobs would be lost at the base in south Wales by June 2013 at the latest.
In October 2010, a £14bn defence training academy which would have created 2,000 jobs at St Athan was scrapped as part of 8% cuts by the UK government to the defence budget.
At that time there were around 400 civilian staff left at the base.
But the operation has been gradually wound down as the contract to service the VC-10 tanker aircraft came to an end.
Mr Lindsay added: "The buildings are there. It's available. Let's hope somebody else comes."
The Welsh government said there was significant private sector interest in the newly designated St Athan Enterprise Zone.
It said it was working to attract new companies to the area capable of creating sustainable jobs for local people.
A spokesman said Cardiff Airport and other key employment sites and development proposals in the immediate vicinity were now being included in the enterprise zone.
"Potential employers are made aware of the skills mix available," said the spokesperson.


Oscar winning musician Annie Lennox, comedian Kevin Bridges and rock band Texas are to perform for former US president Barack Obama when he visits Edinburgh next month.

The dinner on 26 May is thought to be one of his first major addresses since his term as president came to an end.
The event at the EICC is being organised by The Hunter Foundation.
It is being supported by Royal Bank of Scotland and is fully booked.
Mr Obama will also answer questions from the audience at the event, which is being held to raise money for charity.
The foundation was set up by leading businessman and philanthropist Sir Tom Hunter.
All of the cash raised will be split between children's charities in Scotland and the Obama Foundation, set up by the former president and his wife Michelle.
Sir Tom also plans to set aside some seats at the event for local young people.
He said: "Demand for this dinner has been unparalleled in our own experience and we can only apologise to those who we cannot accommodate at the dinner."


Newport Gwent Dragons have signed experienced fly-half Nick Macleod.

The 33-year-old, a former Wales Under-21 international, moves to the region after making more than 150 appearances for Sale Sharks.
"I have strong links with the Dragons through many of the players and coaches, so I am looking forward to my return home," said Macleod.
"It's great to be able to come back to Wales for both personal and professional reasons."
Macleod adds experience to a young Dragons squad.
"The coaches are assembling a young and exciting squad and I am looking forward to joining the region," he said.
"I feel that part of my role next season will be to mentor the youngsters as well as challenging for a spot on the team sheet."
Dragons director of rugby Lyn Jones said: "We have made a strong and conscious decision to back our youngsters and their development, and signing someone of Nick's calibre recognises this as he will act as a catalyst for these players.
"Nick will be mentoring the likes of Dorian Jones, Angus O'Brien and Arwel Robson whilst also playing his part in the squad moving forward next season.
"His vast experience and maturity within the Aviva Premiership will be a valuable addition to our young squad."


US-led forces have stepped up air strikes against Islamic State (IS) fighters threatening the Syrian town of Kobane, near the Turkish border.

The coalition had carried out 21 strikes over two days, a sharp increase that slowed IS advances, the US said.
President Barack Obama predicted a "long-term campaign" against the group, which holds swathes of Syria and Iraq.
He was speaking at a meeting of military commanders from 22 countries that have joined the anti-IS coalition.
"This is an operation that involves the world" against IS, Mr Obama said after the meeting at Andrews Air Force Base near Washington.
"There are going to be periods of progress and setbacks," he added.
President Obama said the allies were "deeply concerned" about the situation in and around Kobane.
The battle for the predominantly Kurdish town has lasted for a month and is regarded as a major test of whether the coalition's air campaign can push back IS.
US-led forces have been carrying out air strikes in support of Kurdish forces there for two weeks.
In a statement, the US military said Tuesday's raids had destroyed IS buildings and military vehicles.
It added that the situation on the ground "remains fluid, with IS attempting to gain territory and Kurdish militia continuing to hold out".
Abdulrahman Kok, a Kurdish journalist in Kobane, told Reuters news agency that the air strikes had lasted throughout Tuesday, calling it "a first".
But he added that IS had intensified its own shelling later in the afternoon.
IS is believed to control about half of the town, from which more than 160,000 people have fled.
Capturing it would give the group unbroken control of a long stretch of the Syrian-Turkish border.
Mr Obama also said the US was "also focused on the fighting that is taking place in Iraq's Anbar province".
The militants have made significant gains in the western province in recent weeks, despite air strikes from the US and its allies since August.
The battles in both Anbar and Kobane illustrated the threat posed by IS and "coalition air strikes will continue in both these areas", he said.
In other developments:


The backlash against Facebook's "free mobile data" Internet.org scheme has spread across the globe.

A total of 67 digital rights groups - including i Freedom Uganda, Ecuador's Usuarios Digitales and Indonesia's ICT Watch - have signed a letter to Facebook's founder, Mark Zuckerberg, stating concerns about the initiative.
They say the project threatens freedom of expression, privacy and the principle of net neutrality.
Facebook has yet to respond.
However, Mr Zuckerberg has previously defended his effort to get people online, by saying: "It's not an equal internet if the majority of people cannot participate."
He has also promised to make changes to address some of the security concerns.
Internet.org allows subscribers of partner mobile networks to use a limited number of online services without having to pay to make use of the data involved.
They include Wikipedia, the Facts for Life health site run by the United Nations Children's Fund, BBC News, Facebook, Accuweather and a selection of local news and sports results providers.
To access the facility, people must use special Android apps, Internet.org's website, Facebook's own Android app or the Opera Mini browser.
The web pages provided must be basic to minimise data use - high resolution photos, videos and voice chat facilities are not permitted.
Network operators participate because they believe users will pay for wider internet access once they have had a chance to try out the free content on offer.
Since 2014, the project has launched in Zambia, India, Colombia, Guatemala, Tanzania, Kenya, Ghana, the Philippines, Indonesia and Malawi.
Facebook says more than nine million people have used the scheme to date.
Until now, the most vocal opposition to Internet.org had come from India's tech community.
Local start-ups complained they risked being disadvantaged because they were not included, while several larger groups that were part of the scheme - including the media conglomerate Times Group and the travel booking site Cleartrip - pulled services, citing concerns about it failing to provide a "fair, level playing field".
The open letter from the 67 digital rights groups - which has been published on Facebook - makes clear that activists across the globe intend to challenge its expansion.
"It is our belief that Facebook is improperly defining net neutrality in public statements and building a walled garden in which the world's poorest people will only be able to access a limited set of insecure websites and services," it states.
"Further, we are deeply concerned that Internet.org has been misleadingly marketed as providing access to the full internet, when in fact it only provides access to a limited number of Internet-connected services that are approved by Facebook and local ISPs [internet service providers].
"In its present conception, Internet.org thereby violates the principles of net neutrality, threatening freedom of expression, equality of opportunity, security, privacy and innovation."
Earlier this month, Mr Zuckerberg sought to address concerns by inviting more organisations to apply to join Internet.org's platform.
But one of his requirements - that the websites should not include the HTTPS, TLS or SSL encryption technologies - caused fresh controversy.
In their letter, the campaigners suggest this will make users' web traffic "vulnerable to malicious attacks and government eavesdropping".
However, a Q&A published on Internet.org's site promises it will begin to support SSL and TLS "in the Internet.org Android app in the coming weeks".
And Mr Zuckerberg has posted a message to his Facebook page promising that support for HTTPS is also under development.
"We still need to do some work to make this work on all phones and browsers - so that's why our docs say it's not currently available - but we're going to make this happen soon," he wrote.
Even so, campaigners say they still have privacy concerns.
"Given the lack of statements to the contrary, it is likely Internet.org collects user data via apps and services," their letter states.
"There is a lack of transparency about how that data is used by Internet.org and its [telecommunications] partners."


The United States shattered a 27-year-old world record to win gold in the women's 4x100m relay at London 2012.

Media playback is not supported on this device
The team of Tianna Madison, Allyson Felix, Bianca Knight and Carmelita Jeter blazed round the track in 40.82 seconds, beating East Germany's record of 41.37, which had stood since 1985.
Jamaica were second in a national record time of 41.41, with Ukraine third in 42.04 and Nigeria fourth.
"That was a fantastic run by the US relay team but the exchanges weren't as crisp as yesterday's heats. They knew by bringing in Allyson Felix and Carmelita Jeter for the final they had speed and I think they decided to go with safer exchanges."
The USA team had failed to reach the final at the Beijing Olympics.
But after qualifying fastest for the 2012 final, they led from the start on a balmy night in east London.
Madison gave the US an electric start, with Felix and Knight maintaining the lead before 100m silver medallist Jeter stormed down the straight to capture the gold.
"I knew that these girls were going to run their hearts out," said Jeter. "I knew we were running fast.
"There was a cloud hanging over us with people saying 'they can't do this, they're going to drop the stick' but we did it."
The Jamaican quartet featured 100m champion Shelly-Ann Fraser-Pryce and bronze medallist Veronica Campbell-Brown, but they could not match the fierce pace of the world champions.
"It's an honour to be part of this team," said Felix, the 200m champion. "Who would have thought that we would have had a world record tonight? It's amazing. Our names are going down in history."
It was the second track world record at London 2012 after Kenya's David Rudisha set a new men's 800m mark on Thursday.


Jurors who found a brother of Gerry Adams guilty of raping his own daughter were not properly directed on how to deal with widespread publicity in the case, the Court of Appeal has heard.

Lawyers for Liam Adams said the level of media coverage on both sides of the Irish border even before his trial turned it into "a national issue".
They also argued guidance to the jury may have wrongly shifted the burden onto him to prove he was innocent of a string of sexual assaults on his daughter Áine. The appeal continues.
Liam Adams, 59, formerly of Bernagh Drive in Belfast, is serving a 16-year jail sentence for the offences against his daughter, who waived her right to anonymity.
The abuse was said to have been committed over a six-year period between 1977 and 1982 when she was aged between four and nine.
He consistently denied the allegations throughout a second trial at Belfast Crown Court in 2013.
However, a jury convicted him of 10 offences against his daughter: three charges of rape, four counts of indecent assault and a further three counts of gross indecency.
At the start of his appeal on Wednesday, he smiled at his wife Bronagh and other relatives who had gathered in the public gallery to support him.
Opening the case, his barrister said they were challenging what she said was the trial judge's failure to direct the jury on how they should assess issues about the extensive publicity before and after a first trial that had collapsed due to legal reasons.
She said a television documentary had sparked widespread media attention.
Her client's brother, Gerry Adams, revealed in an interview that his father subjected family members to sexual abuse, the court heard.
Giving evidence as a prosecution witness at the first trial, the Sinn Féin president claimed his brother confessed to him that he had "molested" his daughter.
Focusing on the level of publicity, the barrister said that by the time of the second trial, any jury member would have heard about the case and her client's earlier battle against being extradited from the Republic of Ireland.
"What makes this case different from a case where someone who is well-known to the public is being tried for serious criminal offences... (is) the complainant, against the advice of police as it turned out, made a television programme," she said.
The barrister also claimed that the trial judge got it wrong in how she directed jurors on dealing with Liam Adams' witness box account.
"It's at the very least open to the danger that the jury may have thought in this case, because the defendant gave evidence, that in some way the burden of proof reverted to him," she said.
The case continues.


A 30cm (1ft) snake slowly moves through the body of a man on a spotless table, advancing its way around the liver.

It stops, sniffs to the left, then turns to the right and slithers behind the ribcage.
This is a medical robot, guided by a skilled surgeon and designed to get to places doctors are unable to reach without opening a patient up.
It is still only a prototype and has not yet been used on real patients - only in the lab.
But its designers, from OC Robotics in Bristol, are convinced that once ready and approved, it could help find and remove tumours.
The mechanical snake is one of several groundbreaking cancer technologies showcased at this week's International Conference on Oncological Engineering at the University of Leeds.
Most of the devices are in very early trial stages, but Safia Danovi from Cancer Research UK says that innovation and research are extremely important in tackling the disease.
"Surgery is a cornerstone treatment for cancer so new technologies making it even more precise and effective are crucial," she says.
"Thanks to research, innovations such as keyhole surgery and robotics are transforming the treatment landscape for cancer patients and this trend needs to continue."
Cancer accounts for about 13% of worldwide deaths every year, according to the World Health Organization.
Although some cancer treatments involve non-invasive methods, surgeons often need to get inside the body - a procedure that is often risky.
Snake robots could be as minimally invasive as possible with today's technology - they would use body orifices or local incisions as points of entry, says Rob Buckingham, managing director at OC Robotics.
They would allow a surgeon to look and "feel" inside the body - by using cameras and extremely sensitive equipment to provide feedback.
Snake robots could complement a robotic surgical system that has been used for the past decade - the Da Vinci machine, developed by US company Intuitive Surgical.
This mechanism looks like a human-size robot with four "arms" equipped with pincers.
Although it cannot perform surgery autonomously, it has allowed doctors to make complex operations less invasive and more precise.
The Da Vinci is controlled by a surgeon sitting in a nearby chair and looking at a screen displaying the area of the body where the surgery is taking place. The surgeon manipulates the robot by pressing pedals and moving levers.
Many hospitals around the world have opted for the Da Vinci, despite the cost of about £1.4m ($2.2m).
Another option is a thin, long mechanical "arm" called Mirosurge, developed by German aerospace centre DLR.
It is also only a prototype, but a DLR engineer who described the technology at the conference says that in the long run, the robot is more versatile than the Da Vinci machine.
"You can attach different tools to it and it can be used either as a one-arm application or as four arms on a surgical table to assist a surgeon controlling them from a workstation," says Sophie Lantermann.
"Also, it has sensors in all the joints, allowing it to detect collisions. For instance, when there's a patient between two arms of the Da Vinci, one arm can hit the other, but our robotic arm detects another arm working next to it.
"It can be a lot cheaper for a hospital because you can use it all day long for different applications."
One of the challenges with cutting away a tumour is making sure that all of it is removed.
For this, the surgeon needs to understand where the tumour stops, which can be tricky.
At the University of Bern in Switzerland, scientists inject a drug into the patient's body and once attached to the tumour, it starts glowing when light is shone on it.
This imaging technology is also applied to instruments as they navigate inside the body, the same way a GPS helps us find our way home.
"It is about tracking surgical instruments, so that a surgeon sees on a computer screen as a medical tool moves through the body, overlaying it with CT scan data - just like an augmented reality application," says Stefan Weber from the ARTORG Center for Biomedical Engineering Research at the University of Bern.
"If you look at the liver, for example, it's a homogeneous, reddish-brownish organ, but you would like to see where the tumours are.
"We scan the patient, derive a 3D model of the liver with the vessels, and in that model we see the tumours so we can tell the surgeon where to cut.
"Detection of vessels, aligning the model with the patient's anatomy, and doing it precisely - even five years ago computers weren't able to do this."
At Leiden University Medical Centre in the Netherlands, scientists are developing a similar technique.
But one of the main topics of discussions at the conference has been about making new technologies work together.
"If we start to combine, for instance, our snake-like robot that can nose-follow to avoid or reach behind organs within the body, with sensors that can identify and lock in on the target that has been marked in some special way, then there may be clinical benefit," says Rob Buckingham.
"Part of the challenge is to think about that benefit. Typically we start travelling in a specific direction, assuming a benefit, and then realise that the goal isn't quite where we expected.
"The important thing is to start moving, to learn."


Queensland recorded their 11th State of Origin series win in 12 years with a thumping victory against New South Wales before their own fans at Suncorp Stadium in Brisbane.

Having lost game one on home turf, Kevin Walters' side fought back to win game two and dominated the decider.
Cronulla winger Valentine Holmes ran in a hat-trick, with Gold Coast prop Jarrod Wallace adding another as the Maroons scored four tries to one.
Josh Dugan got the Blues consolation.
Three series in a row have re-established Queensland's dominance of the Origin series, in spite of the absence of the talismanic Johnathan Thurston through injury.
While the relatively inexperienced Holmes and Wallace picked up the tries and Cameron Munster impressed on his debut, the victory was built on the experience and class of long-serving trio Cameron Smith, Cooper Cronk and Billy Slater.
New South Wales failed to match the Maroons intensity, and had few chances to trouble their rivals on a tough night north of the Tweed River.


Tiger Woods has been named as one of United States captain Davis Love III's assistants for the 2016 Ryder Cup.

Steve Stricker and Jim Furyk will also be joining Tom Lehman as vice-captains for the contest against Europe.
Woods, 39, who is battling to recover from a persistent back injury, has played in seven Ryder Cups, the most recent in 2012 under Love's captaincy.
"Once I'm fully healthy, I'd like to try to make the team too," said 14-time major winner Woods.
"Either way, I'm very excited to work with Davis, the other vice-captains and the players to get a US victory."
Should Woods, now ranked 384 in the world, qualify as a player, either automatically or via a captain's pick, Love has the option to name a replacement vice-captain.
The US will be bidding to end a run of eight defeats by Europe in the last 10 editions of the biennial team competition when the 2016 Ryder Cup is held from 30 September to 2 October at Hazeltine National Golf Club in Minnesota.
Love, who will name a fifth vice-captain at a later date, said: "These three guys, joining Tom Lehman, will send a message to our team that we are really serious about this Ryder Cup.
"We are ahead of the game this time and it lets potential team members know that we are going to do whatever it takes to put them in the best position to win in 2016."


Arlene Foster has said the March election could be the most important since 1998 and would decide the future direction of Northern Ireland.

But the DUP leader said it should not be seen as a referendum on her handling of the Renewable Heat Incentive (RHI) scheme.
She made the comments to the BBC's Inside Politics programme on Friday.
She said Sinn Féin had used RHI as a "useful way to bring the assembly down" and re-run last year's election.
The former first minister claimed republicans wanted to weaken unionism and pursue their wider aims regarding the Irish language, Brexit and increasing the number of cross-border bodies.
She said there had been plenty of occasions when the DUP could have brought the assembly down, but had chosen not to in order to make politics work.
She accused Sinn Féin of walking away from an agreement made between the two parties in mid December on how to handle the controversy over the heating scheme.
Until mid December, Mrs Foster insisted, she and Martin McGuinness had "been working quite well together".
She said it was "absolutely wrong" to characterise her as someone who could not work with Sinn Féin and did not want to make devolution work and that after the election she would have "to try to put it (devolution) back together again".
Mrs Foster repeated her assertion that she had done nothing wrong with regards to RHI and said none of her friends or family "were even remotely connected with the Renewable Heat Incentive scheme, directly or indirectly".
She said rumours to the contrary on social media were incorrect and were "in the hands of my husband's solicitors".
Mrs Foster claimed there was now a "feeding frenzy" around the scheme, and she wishes a public inquiry had been put in place in mid December.
She also said she wishes the cost-cutting measures passed by the assembly earlier this week "had come to the fore sooner".
RHI was also the subject of an exchange between DUP party chairman Lord Morrow and Mrs Foster earlier on Friday.
In response to a comment about it being "a lovely sunny day", Lord Morrow said:  "It's hot. Yes, the boilers are all working."
Mrs Foster replied:  "Stop".


The UK's home secretary has expressed concerns about plans to make it easier for Chinese tourists to get visas.

Culture Secretary Jeremy Hunt had called for the system to be simplified to help "turbocharge" tourism in the wake of the London 2012 Olympics.
But a letter seen by the Daily Telegraph suggests Theresa May believes it would pose a security threat.
The letter, written by Mrs May's private secretary, says she believes the plan is "unacceptable".
On Tuesday, Mr Hunt set out a strategy to give UK tourism a boost following London's Games and said ministers were looking at simplifying visa applications for non-EU nationals.
The strategy focused on attracting visitors from China, where the government thinks there is potential to triple the number of tourists - generating more than Â£500m in extra spending and creating more than 14,000 jobs.
Currently, visitors can apply for a single visa to visit much of Europe - but a separate one is required to travel to the UK.
Countries such as France and Germany are far more successful at attracting Chinese visitors, and the UK could do better if getting a visa was easier, Mr Hunt has suggested.
But a letter from Mrs May's private secretary to the prime minister's private secretary said the current rules were a "key tool in protecting the public against significant harm".
There are already 400 Chinese criminals awaiting deportation and 1,000 asylum applications from Chinese citizens last year, it said.
"The proposal... is not acceptable to the home secretary for national security reasons," the letter said.
"We also face significant challenges with foreign national offenders and organised crime, including drugs, money laundering, fraud, criminal finances, intellectual property, immigration and cyber crime."
But Mark Tanzer, chief executive of the Association of British Travel Agents (Abta), said: "In the absence of concrete data about visa abuse, it is impossible to determine whether the right balance is being struck between the UK's commercial interests and its security needs."
He argued that "proper visa monitoring and reporting processes" were needed, and any abuses of the system could be addressed as they arose.
"We must not forget that visas are only one of the factors that deter Chinese tourists from visiting us - lack of air routes and high aviation taxes also depress demand," he added.
Mr Tanzer has previously said his organisation "welcomed" Mr Hunt's position as "a step in the right direction".
Meanwhile, Immigration Minister Damian Green told that a visa scheme had failed to attract as many of the most talented artists, musicians and scientists to come and work in the UK as the government had hoped.
"We offer a thousand visas a year for the exceptionally talented," Mr Green said.
"In its first year in operation it came nowhere near using them all.
"That tells me we need to fine tune the system to make sure that around the world, if you are a bright young artist or a gifted young scientist, you know there is this route to come and work in Britain. We need to do better on that."
The minister added: "If you are a current Nobel Prize winner, you won't find it very difficult to get into this country.
"But we want the next generation of Nobel Prize winners, so we have carved out a specific route in the immigration system."


Mark Selby completed the perfect preparation for his world title defence, beating fellow Englishman Gary Wilson in the final of the China Open.

Selby, 31, made a break of 145 in frame seven as he won 10-2 in Beijing.
Former taxi driver Wilson, ranked 56th, had shocked defending champion Ding Junhui 6-5 in Saturday's semi-finals.
The World Championship gets under way in Sheffield on 18 April, with Selby hoping to match last year's feat when he beat Ronnie O'Sullivan in the final.
"I played well for the majority of the match," Selby told World Snooker.
"To be 8-1 up at the end of the first session probably wasn't a fair reflection on how the match had gone, as a few frames could have gone either way but I managed to pinch them.
"I'll get home tomorrow, have a few days off and then get back on the practice table to prepare for the Crucible."
Selby moved 3-0 clear in Sunday's Beijing final before Wilson briefly slowed his progress with a break of 91.
The world number one showed he was in top form when the highest break of the tournament at 145 took him 6-1 clear, and a 136 clearance made it 9-1.
Wilson had never previously been beyond the quarter-finals of a ranking event, but the 29-year-old from Wallsend gained some consolation with a break of 101 to get on the board for a second time.
Selby was not about to be knocked off course, however, and wrapped up victory with seven frames to spare to earn his second ranking title of the season.
"I'm disappointed to have played like that and lost by such a heavy score line," said Wilson, who collected his biggest career cheque of £35,000.
"I missed a lot of chances. I can still take some positives from the tournament and look forward to the World Championship."


Mae llywydd Ymddiriedolaeth Cefnogwyr clwb pêl-droed Wrecsam, Bryn Law, wedi camu i lawr ac wedi cwestiynu perchnogaeth y cefnogwyr o'r clwb.

Daw penderfyniad Mr Law yn dilyn ymddiswyddiad y prif weithredwr, Don Bircham a chyfarwyddwr y clwb, Barry Horne gyda'r clwb yn 15 safle yn y Gyngres.
Mae Ymddiriedolaeth y cefnogwyr yn berchen ar 100% o'r clwb ond dywedodd Law ei fod yn parhau i gefnogi "delfryd yr Ymddiriedolaeth ond rwyf yn dechrau meddwl tybed a yw'n gweithio yn y ffordd orau i'r clwb."
Roedd Mr Law yn llywydd ymddiriedolaeth y cefnogwyr ers 2010.


Lebanon is holding a day of national mourning after at least 41 people were killed in two suicide bombings in the capital, Beirut.

The Islamic State (IS) militant group says it was behind the attacks in Burj al-Barajneh, a mainly Shia southern suburb and Hezbollah stronghold.
Hezbollah forces are fighting IS in neighbouring Syria.
The bombing is the deadliest in the capital since the end of Lebanon's civil war in 1990.
Lebanese Prime Minister Tammam Salam called for unity in the face of attempts to create strife.
In reaction from Washington, the White House in a statement condemned the "horrific terrorist" bomb blasts which it said "would only serve to reinforce our commitment to support the institutions of the Lebanese state, including the security services, to ensure a stable, sovereign and secure Lebanon".
Iran - a key ally of Hezbollah - has also condemned the attack.
Lebanese media denounce blasts
The blasts on Thursday evening struck in a busy shopping street. More than 200 people were wounded - many of them seriously, Health Minister Wael Abou Faour said.
The army said that two men wearing suicide vests carried out the attack. The first bomber detonated his explosive vest outside a Shia mosque, while the second blew himself up inside a nearby bakery.
The body of a third bomber who failed to detonate his explosives was found at the scene of the second blast.
An IS statement - which could not be independently verified - said that its "soldiers of the Caliphate" detonated explosives which were concealed on a motorbike.  It identified the three attackers as two Palestinians and a Syrian.
"After the apostates gathered in the area, one of the knights of martyrdom detonated his explosive belt in the midst of them," the statement said. It did not refer to Hezbollah's involvement in Syria.
Hezbollah has in turn accused IS of targeting innocent civilians, including unarmed people and the elderly. "It was a Satanic terrorist attack," Hezbollah official Bilal Farhat told the Associated Press.
Hezbollah has vowed to continue its fight against "terrorists", warning of a "long war" against its enemies, according to the Reuters news agency.
BBC Arab affairs analyst Sebastian Usher says it is no surprise that IS has claimed these bombings, but their ferocity will once again reawaken the spectre of Lebanon's 15-year civil war.
Hospitals in southern Beirut have urged people to donate blood and called on residents not to congregate at hospital gates so that ambulances and emergency staff could get in and out without being delayed.
Witnesses described a hellish scene in the aftermath of the blasts which took place minutes apart from each other.
"I'd just arrived at the shops when the blast went off. I carried four  bodies with my own hands, three women and a man, a friend of mine," a man who identified himself as Zein al-Abideen Khaddam told local television.
Another described the deafening sound: "When the second blast went  off, I thought the world had ended."
US Secretary of State John Kerry has meanwhile urged countries attending Syrian peace talks this weekend to show flexibility - he said that while diplomatic efforts to bring the civil war to an end were not on the verge of success, all parties in the conflict had a responsibility "not to dig in their heels so that the bleeding can stop".
Prime Minister Salam condemned the attacks as "unjustifiable" and called on Lebanon's rival factions to unite against "plans to create strife".
Hezbollah's strongholds in southern Beirut were the target of a series of bombings in 2013 and 2014 mostly claimed by Sunni jihadist militants - who denounced Hezbollah's decision to send fighters to neighbouring Syria to prop up President Bashar al-Assad.
In recent weeks, Hezbollah has sent reinforcements to Syria in support of government offensives in northern areas held by rebel forces or IS.


A report to prevent future deaths was issued after a woman died following discharge from Wrexham Maelor Hospital.

Carol Harvey died, aged 74, in April last year due to the effects of paracetamol overdose and a pre-existing heart condition, an inquest heard.
Coroner John Gittins recorded a narrative conclusion, saying she was probably left in "considerable pain".
He outlined concerns to Betsi Cadwaladr University Health Board, which apologised to Mrs Harvey's family.
Mrs Harvey was treated at the hospital but was discharged on 7 April and was later found drowsy and vomiting having suffered a large loss of blood.
The north Wales coroner said a message about her care left for a health team was not checked which meant carers did not visit.
This meant "it is probable that in the course of that evening and overnight she was in considerable pain as a result of a calf haematoma and consequently took an accidental overdose of painkillers", the report said.
After being re-admitted to hospital, she died on 9 April.
Mr Gittins has asked the health board for a detailed response and a timetable for action within 56 days.
The health board's medical director, Dr Evan Moore, said: "We apologise to Mrs Harvey's family for the missed opportunities to provide further care, and fully accept the coroner's findings.
"We have since put in place new measures to enhance communication between staff in different wards and departments, as well as a review of the way we discharge patients from acute hospitals on anticoagulant therapy."


David Natzler has been named as the next Clerk of the Commons, ending a controversial process which pitted Speaker John Bercow against some MPs.

Mr Natzler has served as acting Clerk since the retirement of Sir Robert Rogers last year.
A panel led by Mr Bercow initially recommended that Australian Carole Mills take the role but MPs objected and she has since withdrawn.
News of Mr Natzler's appointment was met with cheers by MPs in the Commons.
The recruitment of a new Clerk, Parliament's most senior legal and constitutional adviser, has been dogged by argument over the past nine months.
MPs said Ms Mills, the original choice of the House of Commons commission who works in the Australian Parliament, did not have the relevant constitutional experience and accused Mr Bercow of a "stitch-up".
Mr Bercow insisted that the appointment must have the confidence of MPs and "paused" the recruitment process before re-starting it.
As part of a modernisation of procedures, the current responsibilities of the Clerk will now be split, with a new post of Commons director-general taking over responsibility for the day-to-day commercial and administrative functions of the Commons.
Announcing Mr Natzler's appointment in the Commons, Mr Bercow said he had been chosen from a field of four candidates.
Welcoming the appointment, David Cameron joked that Mr Bercow had "gone to the ends of the earth" to find the most qualified candidate and he was "very glad he found the right answer right here in Britain".
Labour leader Ed Miliband said Mr Natzler's appointment was "well deserved".
Before Sir Robert's retirement, Mr Natzler served as Clerk Assistant for many years.


Thousands more teachers will be needed to work as examiners as qualification reforms kick in, suggests a report.

About 34,000 examiners currently set and mark eight million GCSEs and A-levels for two million 15-19-year-olds in England, Wales and Northern Ireland.
But changes to qualifications and a rise in student numbers mean about 20% more will be needed by 2019, says a panel of exam boards and head teachers.
The report heralds a push to recruit more teachers as examiners.
Reforms to exams are under way in all three nations which are, according to the report, "creating new challenges for the system".
The report highlights changes to qualifications in England which "are driving a change" in the numbers of examiners needed.
In particular, cuts to the amount of coursework in England will see the hours of exam assessment in most GCSEs increase from two-and-a-half to three-and-a-half, say the authors
Additionally, the reformed exams require less knowledge recall from candidates and more analysis and so more of them will "need to be marked by a highly skilled subject expert", they add.
The panel, which includes representatives from the major exam boards as well as head teachers' leaders from both the public and private sectors, also suggests that fewer resits under the new system will reduce the overall number of exams - but will add to demand for examiners during the peak summer marking period.
Finding enough teachers to become examiners could be particularly difficult in subjects where there are shortages, says the report.
And teachers who are struggling with their daily workload may not want to take on the extra burden of becoming examiners, it adds.
In addition, the pay structure for examiners which is often per script or per question can put off teachers, making examining feel like an administrative job rather than "one for skilled and experienced professionals", say the authors.
"It can also make it difficult for prospective examiners to understand how much they will be paid and to value their time appropriately."
In a bid to boost examiner recruitment the Joint Council for Qualifications, representing exam boards has launched:
In addition head teachers' organisations agreed to encourage schools and colleges to promote examining to staff, provide practical support and to recognise examining as a way of boosting teachers' subject knowledge and professional skills.
"The nature of the reforms is that we will need more examiners... and we want to ensure that the quality of these examiners is good," said Dale Bassett, head of curriculum strategy at the AQA board and chairman of the working group which produced the report.
"Rather than waiting for this to become a problem we are proactively working to recruit these extra examiners," Mr Bassett added.
School Standards Minister Nick Gibb said: "It is important that exams are marked to a high quality and standard.
"We welcome the work that the Joint Council for Qualifications (JCQ), awarding organisations, and head teacher associations are doing to recognise the valuable role of examiners."


Thomas Starzl, the man who performed the world's first liver transplant, has died days short of his 91st birthday.

The American surgeon pioneered the procedure in 1963, but his first patient did not survive.
After creating a new blend of anti-rejection drugs, he carried out the first successful transplant in 1967. Since then, thousands of lives have been saved by the procedure.
He died at home among his family, a spokesperson said.
In a statement, the University of Pittsburgh, which he joined in the 1980s to work on his drugs research, said Dr Starzl was known as the "father of transplantation" for his work in advancing the surgery from "from a risky, rare procedure to an accessible" one.
In addition to performing the first successful liver transplants, he experimented with transplants from cadavers, and refined the process by using identical twins and blood relatives.
He also pioneered animal-to-human liver transplants, including baboon to human experiments, which he showed could briefly extend life when there was a shortage of human organs.
His family issued a statement saying he "brought life and hope to countless patients".
"He was a pioneer, a legend, a great human, and a great humanitarian," it said.
"He was a force of nature that swept all those around him into his orbit, challenging those that surrounded him to strive to match his superhuman feats of focus, will and compassion."
Dr Starzl was also known for his research work on developing anti-rejection drugs. He blended azathioprine, a drug which suppresses the immune system, with steroids to aid in his pioneering transplants in the 1960s.
His research later in life would lead to the acceptance of improved drugs including cyclosporine and tacrolimus.
He retired from clinical work in 1991 and published his autobiography, The Puzzle People.
In it, he revealed that despite all his accolades, he felt a great anxiety about actually performing surgery.
"I had an intense fear of of failing the patients who had placed their health or life in my hands," he wrote.
"Even for simple operations I would review books.... then, sick with apprehension, I would go to the operating room, almost unable to function until the case began."
"Instead of blotting out the failures, I remembered these forever," he said.


Voting to leave the European Union was a "bad decision" for the UK, the Irish foreign minister has said.

Charlie Flanagan said Brexit could be "painful" for the UK and his country, which he added should not be "placed in a position of more disadvantage".
He also urged negotiators to keep the Common Travel Area between Northern Ireland and the Republic.
The comments come after Prime Minister Theresa May met European Council President Donald Tusk in London.
Mr Flanagan told BBC's Newsnight it was "absolutely essential" there was no return to a hard border on the island of Ireland - which is expected to be a key element of Brexit negotiations between the UK and EU.
"The Good Friday Agreement remains the foundation stone of our peace, and anything adverse to that agreement will not be acceptable," he said.
However, despite his criticism of the decision to leave, the Irish minister said he believed there was no intent to punish Britain among EU members, adding that the relationship between the UK and the Republic of Ireland was its "warmest ever".
He said: "I believe [Brexit] was a bad decision, but of course as a democrat I fully respect and recognise the will and wishes of the British people. We've got to deal with that now.
"The Article 50 process has commenced, and I believe it's essential now that we get through the negotiations in such a way that the end result can be as close as possible a relationship between the European Union and the United Kingdom, albeit with the UK gone."
On Friday, Mrs May and Mr Tusk met at 10 Downing Street for the first time since Article 50 was triggered, with Gibraltar at the heart of the talks.
In its draft negotiating guidelines, published last week, the EU said decisions affecting Gibraltar - a UK overseas territory - would have to be taken with the agreement of the Spanish government.
This led to former Conservative leader Lord Howard saying the prime minister would defend Gibraltar in the same way Margaret Thatcher defended the Falkland Islands.
Mrs May laughed off the prospect of war with Spain, but after her meeting with Mr Tusk, a Number 10 spokesman said she had been clear she was determined to achieve the "best possible deal" for Gibraltar as well as the UK.
"The PM also made clear that... there would be no negotiation on the sovereignty of Gibraltar without the consent of its people," he added.
EU sources told the Press Association it had been a "good and friendly" meeting.
One source added: "They agreed to stay in regular contact throughout the Brexit process to keep a constructive approach and seek to lower tensions that may arise, also when talks on some issues like Gibraltar inevitably will become difficult."


Paralympic silver and bronze medallist Charlotte Henshaw has retired from swimming and taken up canoeing.

The Briton, who won silver at London 2012 and bronze at Rio 2016, holds the SB6 100m breaststroke European record.
But, after being identified by British Canoeing, the 31-year-old is a member of their talent confirmation programme.
"I watched the brilliant Paracanoe performances in Rio and was keen to try something new," said Henshaw, who has had both legs amputated above the knee.
"I felt like I was ready to take on a new challenge that is different from anything I have done before."
Henshaw began canoeing at the end of 2016 and hopes to qualify for the European Championships as she sets her sights on competing at another Paralympics.
"A fourth Paralympic Games would be beyond what I had ever imagined, but I would love to be there in Tokyo - on the water instead of in it this time," she added.


A man has said a bird of prey caused two long cuts across his head during an attack while he was out jogging.

Andy Llewellyn was running down Infinity Park Way, near Chellaston Medical Centre, Derby, on Tuesday when he felt an "almighty slap" on his head.
He said his wound was treated by medical staff and he was given antibiotics.
The RSPB said it was rare that a bird is so "aggressive" but it was likely to have been protecting its chicks.
More on this and more stories from the East Midlands
Mr Llewellyn said he didn't see the bird but "heard the flapping of wings on my shoulder as it hit my head".
"I think I was a little bit dazed for a few moments. I put my hand on there and I had lots of blood," he added.
"I got home and went to the [NHS] walk-in centre and they said they had never seen anything like it.
"It's made me think a little bit more about the route I take... The ironic thing after the attack is that I did make my fastest time."
An RSPB spokesman said it was not possible to say for certain what type of bird it was, but added that such aggressive behaviour was "rare".
He said: "While clearly a distressing incident for the person involved, this is not usual behaviour.
"The jogger was in all probability running close to a nest, and as we are coming to the end of the breeding season when chicks are starting to fledge, the reaction of the bird was one of a protective parent."


England opened up a 489-run lead over Pakistan after choosing not to enforce the follow-on in the second Test.

Media playback is not supported on this device
The hosts reached 98-1 on the third day at Old Trafford, building on a first-innings lead of 391 after bowling Pakistan out for 198 thanks to Chris Woakes' 4-67.
Captain Alastair Cook is unbeaten on 49 and Joe Root 23, after Alex Hales fell to Mohammad Amir for 24.
No team has chased more than 418 in the fourth innings to win a Test.
Pakistan, 57-4 overnight, slipped to 119-8 before skipper Misbah-ul-Haq (52) and Wahab Riaz (39) added 60 for the ninth wicket.
They can expect to bat again early on the fourth day, as England press for the victory that would level the four-match series at 1-1.
Media playback is not supported on this device
Only 60.4 overs were bowled on a day when there were four interruptions for rain, including two during England's second innings in bowler-friendly conditions.
Former England captain Michael Vaughan said he was "absolutely staggered" by Cook's decision to bat again.
"They were 391 runs ahead - 391 runs! Win the game today - why delay it?" he said on BBC Test Match Special.
"Somebody has tweeted me to say England's sports science department want to manage the bowlers' workload.
"They've only bowled 60 overs. I know I sound like Geoffrey Boycott, but come on!"
Only two teams have won a Test after following on, with Australia beaten by England in 1894 and 1981 and by India in 2001.
Former England batsman Boycott said on TMS: "I can't help but think England have made a mistake.
"If they had enforced the follow-on, it would have been perfect because they've been on and off and the bowlers would have had a rest."
England assistant coach Paul Farbrace said it was a "pretty simple decision" not to enforce the follow-on.
"We wanted to get our runs while it's still a very good pitch, at no point putting ourselves under undue pressure, trying to chase a score. The pitch will deteriorate.
"We've got two days to bowl them out, we can bat on in the morning and get a few more then look to put them under extreme pressure."
Asked if England's decision not to enforce the follow-on was to help protect seamers James Anderson and Ben Stokes, both of whom are returning from injury after missing the first Test, Farbrace said: "No, they're both fully fit."
Pakistan coach Mickey Arthur said: "I fully expected us to be batting again, as did all our batters, who were almost padding up.
"We were probably a bit relieved that we weren't straight back out there. Our batters certainly were happy."
Pakistan, who lost four wickets in 24 overs on the second evening, fared marginally better on Sunday, with only Misbah offering sustained resistance on a pitch that remains reliable.
James Anderson, on his home ground, removed Shan Masood for 39 courtesy of an edge to Root at second slip.
Asad Shafiq drove Stuart Broad loosely to backward point, Sarfraz Ahmed edged Ben Stokes to second slip to fall for 26 off 18 balls, and Yasir Shah followed one from Woakes to give Root, England's double centurion, his fourth catch.
Struck on the helmet by a Woakes bouncer, Misbah received valuable support from Wahab, who hit Moeen Ali for six during the highest partnership of the Pakistan innings.
Misbah, in attempting to dominate Moeen - as he did in the first innings at Lord's - top-edged a sweep to Cook at short fine-leg.
Wahab, who was hit on the left elbow by Woakes and did not bowl in England's second innings, swung Moeen to Hales at deep mid-wicket to end the innings.
Boycott said: "Pakistan almost couldn't resist giving their wickets away.
"You would have thought this morning they would think about it and say 'let's show some fight'."
Media playback is not supported on this device
BBC cricket correspondent Jonathan Agnew
If it had not been for that ninth-wicket stand of 60 between Misbah and Wahab, England would probably have put Pakistan in again.
Not enforcing the follow-on will be viewed as a negative decision but it's not totally illogical as James Anderson and Ben Stokes have just returned from injuries.
It's a statement from the team on top as much as anything else, because whenever Pakistan bat again they will always have a huge target hanging over them and know they have no chance of reaching it.
But I cannot help but think that, had England bowled again, they would have done very nicely in these conditions.


Cambridge United twice came from behind to secure a thrilling extra-time win over National League Dover Athletic and a place in the FA Cup second round.

Media playback is not supported on this device
Ricky Miller's calm one-on-one finish seemed set to earn Dover a shock replay victory over their League Two visitors.
However, Mark Roberts shot home in the 90th minute to force the extra period.
A stunning Aswad Thomas volley restored the hosts' lead, but Leon Legge levelled and late goals by Medy Elito and Ben Williamson sealed victory.
Defender Legge converted a Roberts cross with 10 minutes remaining to make it 2-2 and, after Elito fired in a third away goal, Williamson's solo effort sealed a second-round tie against Coventry in the 121st minute of a pulsating match.
Match ends, Dover Athletic 2, Cambridge United 4.
Second Half Extra Time ends, Dover Athletic 2, Cambridge United 4.
Goal!  Dover Athletic 2, Cambridge United 4. Ben Williamson (Cambridge United) right footed shot from the right side of the box to the centre of the goal. Assisted by Uche Ikpeazu.
Substitution, Cambridge United. Josh Coulson replaces Piero Mingoia.
Goal!  Dover Athletic 2, Cambridge United 3. Medy Elito (Cambridge United) left footed shot from the left side of the six yard box to the bottom left corner.
Attempt missed. Luke Berry (Cambridge United) right footed shot from outside the box is high and wide to the right from a direct free kick.
Uche Ikpeazu (Cambridge United) wins a free kick in the attacking half.
Foul by Jamie Grimes (Dover Athletic).
Attempt saved. Mitchell Pinnock (Dover Athletic) left footed shot from outside the box is saved in the top centre of the goal.
James Dunne (Cambridge United) is shown the yellow card for a bad foul.
Foul by James Dunne (Cambridge United).
Loui Fazackerley (Dover Athletic) wins a free kick in the attacking half.
Foul by Uche Ikpeazu (Cambridge United).
Sam Magri (Dover Athletic) wins a free kick in the defensive half.
Attempt saved. Ricky Modeste (Dover Athletic) right footed shot from a difficult angle and long range on the left is saved in the top left corner. Assisted by Ross Lafayette.
Goal!  Dover Athletic 2, Cambridge United 2. Leon Legge (Cambridge United) right footed shot from the centre of the box to the high centre of the goal. Assisted by Mark Roberts with a cross.
Corner,  Cambridge United. Conceded by Richard Orlu.
Uche Ikpeazu (Cambridge United) wins a free kick on the left wing.
Foul by Sam Magri (Dover Athletic).
Attempt missed. Ross Lafayette (Dover Athletic) left footed shot from outside the box is close, but misses to the left. Assisted by Mitchell Walker.
Attempt saved. Uche Ikpeazu (Cambridge United) right footed shot from the centre of the box is saved in the bottom left corner. Assisted by Piero Mingoia.
Second Half Extra Time begins Dover Athletic 2, Cambridge United 1.
First Half Extra Time ends, Dover Athletic 2, Cambridge United 1.
Attempt missed. Uche Ikpeazu (Cambridge United) left footed shot from outside the box is close, but misses to the right. Assisted by James Dunne.
Foul by Brad Halliday (Cambridge United).
Ricky Modeste (Dover Athletic) wins a free kick in the defensive half.
Substitution, Cambridge United. Blair Adams replaces Greg Taylor because of an injury.
Goal!  Dover Athletic 2, Cambridge United 1. Aswad Thomas (Dover Athletic) left footed shot from outside the box to the bottom right corner following a corner.
Corner,  Dover Athletic. Conceded by Piero Mingoia.
Attempt saved. Uche Ikpeazu (Cambridge United) right footed shot from outside the box is saved in the centre of the goal. Assisted by Greg Taylor.
Attempt blocked. Aswad Thomas (Dover Athletic) right footed shot from outside the box is blocked. Assisted by Ross Lafayette.
Attempt missed. Loui Fazackerley (Dover Athletic) right footed shot from outside the box misses to the left.
Attempt missed. Ross Lafayette (Dover Athletic) header from the right side of the six yard box misses to the left. Assisted by Mitchell Pinnock.
Attempt missed. Luke Berry (Cambridge United) header from the centre of the box is close, but misses to the left. Assisted by Piero Mingoia with a cross.
Attempt missed. Leon Legge (Cambridge United) header from the centre of the box is close, but misses to the left. Assisted by Piero Mingoia with a cross.
Hand ball by Aswad Thomas (Dover Athletic).
Ben Williamson (Cambridge United) wins a free kick in the defensive half.
Foul by Loui Fazackerley (Dover Athletic).
Corner,  Dover Athletic. Conceded by Greg Taylor.
Foul by Leon Legge (Cambridge United).


All week on Newsround we've been talking about the issue of bullying, something that affects many people.

20 February 2015 Last updated at 17:46 GMT
It's an important issue and it's vital that the subject is addressed to try to find out how to deal with it.
Jenny went to a Diana Award Anti-Bullying Ambassadors event at the Emirates Stadium, where some celebrities were running workshops for children on how to tackle bullies.
Hear from the likes YouTube stars Niki and Sammy, and musical duo Bars and Melody - plus a few other familiar faces!
Click here for a preview of the new Newsround special, How to Make Friends on Planet Earth.


The roar of shareholder disapproval became deafening yesterday as Weir Group became only the second company in UK history to lose a binding vote on executive pay.

We have seen other insurrections - most notably at BP - but these have been non binding. What's the difference?
Shareholders get to vote on pay every year but only once every three years does their vote have any teeth.
This is the moment when the company presents the pay policy. In other words when it lays out the method by which pay will be calculated for the next three years. If shareholders reject this - as Weir's did yesterday - its back to the drawing board.
The debate over high pay tends to focus on the injury done to our ideas of fairness and merit. The gap between the boardroom and the shop floor has become a chasm.
According to the High Pay Centre, 20 years ago CEOs earned 40 times as much as the average worker. That multiple is now 180.
There is increasing support gathering for the view that high pay is not just grossly unfair and socially divisive, it is economically destructive.
How so? The argument goes like this.
Remember, most company CEOs don't last long. The average tenure for a FTSE 100 boss is five years. Even when bonuses and pay are delayed, the period over which their performance is judged is pretty short.
Therefore, the CEO has to work quickly. The best way to maximise profit is to keep prices as high as the market will bear while cutting costs to the bone.
That will increase profits in the short term but is also a recipe for a lack of investment in new products, new machinery, new training, new technology, new anything.
All the things that, over time, raise an economy's productivity and its citizens living standards. The amount of money businesses are investing in things that will provide long term benefits is lower than at any time since the 1960s.
Some writers, like economist Andrew Smithers, argue that it is not too far fetched to link the decline in productivity, the amount of output per worker hour, to the excesses of executive pay.
We have been here before. The shareholder spring of 2012 was meant to chasten boards into more comprehensible, more transparent, and less offensive pay awards. It ushered in the idea of a three-yearly binding vote that Weir just lost.
And yet here we are today with Bob Dudley trousering £14m as his shareholders suffer and, wait for it, Sir Martin Sorrell to get a pay award of nearly £70m (his shareholders made money).
Senior city figures realise something is wrong. Nigel Wilson, the chief executive of Legal and General last week launched a working group to examine a pay regime he described as "broken".
It remains to be seen whether this shareholder spring gives way to yet another long summer for those in the boardroom.


Gloucestershire finished day one on 62-3 in reply to Glamorgan's 220 all out at the SSE Swalec Stadium in Cardiff.

Glamorgan's total was dominated by Nick Selman's mature 101 off 144 balls, his second first-class century.
But seamers Matt Taylor and Craig Miles made sure wickets fell at regular intervals in cloudy conditions in the Welsh capital.
Gloucestershire struggled in turn as the bowlers remained on top all day.
Selman's century, after being dropped on eight in the first over, was a welcome return to form for the Brisbane-born opener after a string of low scores following his hundred against Northants at Swansea.
Aneurin Donald's quick-fire 36 off 30 balls was the next best contribution as Taylor and Miles prospered in the afternoon.
Graham Wagg and Timm van der Gugten scrambled Glamorgan to a consolation bonus point.
Gloucestershire's innings was a question of survival in the evening gloom, with the day ending evenly balanced.
Glamorgan are being led by Michael Hogan for the first time in the absence of Jacques Rudolph, who has a neck problem.
Glamorgan batsman Nick Selman told BBC Wales Sport:
"It was good to get some runs for a change, the wicket was a bit slow and they bowled a bit short early on so I got through the first 25 overs and it got easier to bat from there, but it was a bit disappointing to get out just after I got a hundred.
"(My season) was going well up until after (a century in) Swansea, after Swansea four ducks in a row, but it was good to spend some time in the middle at Northants.
"Opening the batting in county cricket over here (in the UK), it's definitely hard, you get some good balls, you play on some green wickets and I was struggling a bit.
"I was happy to get a few today and hopefully I can continue that for the rest of the season."
Gloucestershire seamer Matt Taylor told BBC Radio Bristol:
"There's definitely a bit on offer out there (for the bowlers), we didn't start exactly how we'd like but we got there in the end and we'd take bowling them out for 220.
"I was pretty happy with that (hitting the stumps three times), the same plan as always is to hit the top of off-stump and luckily they missed them today.
"I was pleased to get an opportunity in T20 and the white-ball stuff this year, now I'm getting an opportunity with the red ball it's great.
"It's definitely going to be tough (for the Gloucestershire batsmen), they're a good bowling attack but hopefully we can dig in and get to a reasonable score."


A 12-year-old German-Iraqi boy tried to blow up a Christmas market in the town of Ludwigshafen, prosecutors say.

He left a rucksack with explosives which failed to detonate in November, and put another explosive device by the town hall a few days later, they say.
But a member of the public alerted the police, and no-one was injured.
The 12 year old - who is now in custody - is thought have been recently radicalised and to have links with the so-called Islamic State group.
Focus magazine cited security and judicial sources as saying the boy was "strongly radicalised" and apparently instructed by an unknown IS member.
The Federal Public Prosecutor's Office declined to comment on a possible IS link, but confirmed officials were investigating the case.
Public Prosecutor Hubert Stroeber said the boy left a backpack containing a home-made nail bomb at the Christmas market in Ludwigshafen, in the state of Rhineland-Palatinate, on 26 November, but the device did not go off because the detonator apparently failed.
The second device near the city's town hall, placed on 5 December, was discovered by an "informant"  and specialists defused it, Mr Stroeber said.
The boy was born in Ludwigshafen to Iraqi parents in 2004.
Prosecutors said no formal proceedings would be launched against him, because he is below the age of criminal responsibility.
Last Thursday, German police arrested two teenagers aged 15 and 17, accused of plotting a terror attack on a public institution in Aschaffenburg, in north-western Bavaria. Police found Islamic State flags and propaganda material in their homes.
Germany suffered a spate of attacks in the south of the country over the summer.
Ten people were killed and dozens more injured in separate gun, bomb, axe and machete attacks.
The authorities say they were not linked, and one of them was not terror-related.
But the wave of violence has made a nation already on edge after a huge influx of refugees even more nervous.


Ryan Blake's strike sent Southern League Premier Division side Chesham into round two of the FA Cup at the expense of League Two Bristol Rovers.

Media playback is not supported on this device
Chesham should have been ahead at half-time after Tom Lockyer fouled Brad Wadkins, but Rovers keeper Lee Nicholls saved Dave Pearce's resulting penalty.
But the seventh tier side did take the lead when Blake's powerful shot from 25 yards caught out Nicholls.
Shane Gore made a string of saves late on as Chesham held on.
Ellis Harrison and Jermaine Easter were among those thwarted by the visitors' keeper as Rovers applied late pressure in the search for an equaliser.
Chesham, who sit 75 places below Rovers in the football pyramid, last made the first round of the FA Cup in 1994.
Victory sees the Generals progress to the second round for only the second time in their history, the first coming in 1980 when they were knocked out in the third round by Cambridge United.
Former Rovers striker and current Chesham player-coach Barry Hayles was given a warm ovation when he appeared as a 72nd-minute substitute, minutes before he supplied the assist for Blake's winner.
Match ends, Bristol Rovers 0, Chesham United 1.
Second Half ends, Bristol Rovers 0, Chesham United 1.
Foul by Ellis Harrison (Bristol Rovers).
George Casey (Chesham United) wins a free kick in the attacking half.
Corner,  Bristol Rovers. Conceded by Bradley Wadkins.
Corner,  Bristol Rovers. Conceded by Stephen Hamilton-Forbes.
Corner,  Bristol Rovers. Conceded by Bradley Wadkins.
Substitution, Chesham United. Stephen Hamilton-Forbes replaces Matt Taylor because of an injury.
Attempt saved. Jake Gosling (Bristol Rovers) right footed shot from outside the box is saved in the top centre of the goal.
Corner,  Bristol Rovers. Conceded by Toby Little.
Attempt saved. Jermaine Easter (Bristol Rovers) right footed shot from the centre of the box is saved in the centre of the goal.
Foul by Chris Lines (Bristol Rovers).
Bradley Wadkins (Chesham United) wins a free kick in the attacking half.
James Clarke (Bristol Rovers) wins a free kick in the attacking half.
Foul by Barry Hayles (Chesham United).
Foul by Tom Lockyer (Bristol Rovers).
Ashley Smith (Chesham United) wins a free kick in the attacking half.
Corner,  Bristol Rovers. Conceded by Shane Gore.
Attempt saved. Ellis Harrison (Bristol Rovers) left footed shot from the centre of the box is saved in the centre of the goal.
Substitution, Chesham United. George Casey replaces Ryan Blake because of an injury.
Corner,  Bristol Rovers. Conceded by Barry Hayles.
Corner,  Bristol Rovers. Conceded by Ashley Smith.
Corner,  Bristol Rovers. Conceded by Curtis Ujah.
Attempt blocked. Jermaine Easter (Bristol Rovers) left footed shot from the right side of the box is blocked.
Foul by Chris Lines (Bristol Rovers).
Ryan Blake (Chesham United) wins a free kick on the left wing.
Attempt missed. Tom Lockyer (Bristol Rovers) header from the centre of the box misses to the right following a corner.
Corner,  Bristol Rovers. Conceded by Nick Beasant.
Attempt blocked. Jermaine Easter (Bristol Rovers) right footed shot from the centre of the box is blocked.
Goal!  Bristol Rovers 0, Chesham United 1. Ryan Blake (Chesham United) right footed shot from outside the box to the bottom left corner.
Attempt blocked. Billy Bodin (Bristol Rovers) right footed shot from outside the box is blocked.
Corner,  Bristol Rovers. Conceded by Ryan Blake.
Foul by Jake Gosling (Bristol Rovers).
Bradley Wadkins (Chesham United) wins a free kick on the left wing.
Substitution, Bristol Rovers. James Clarke replaces Daniel Leadbitter because of an injury.
Substitution, Chesham United. Barry Hayles replaces David Pearce.
Toby Little (Chesham United) is shown the yellow card for a bad foul.
Billy Bodin (Bristol Rovers) wins a free kick in the defensive half.
Foul by Toby Little (Chesham United).
Stuart Sinclair (Bristol Rovers) wins a free kick on the right wing.


A P&O ship steward charged with conspiring to import cocaine into Hull on a ferry has been accused of lying to try to "save his own skin".

Edward Tron, 51, of Carr Hill Road, Gateshead, claims stories about drug smuggling he told an undercover police officer were fabricated.
At Hull Crown Court, the prosecution alleged Mr Tron was attempting to "hide the position" he was in.
Mark Quilliam, 55, of Gladica Close, Liverpool, faces the same charges.
More on this and other Hull stories
Mr Tron's wife Susan, 54, also of Carr Hill Road, is accused of money laundering, with all three denying the charges against them.
The court previously heard Mr Quilliam and Mr Tron used their positions as P&O ferry crew to smuggle cocaine into Hull from Rotterdam on the Pride of Hull vessel.
It is claimed they made up to Â£60,000 a trip.
Mr Tron, who told police he had made Â£100,000 from gambling, told the court his gambling started to "hit lucky for four years" before his luck ran out.
"I thought I was invincible at one time", he said, but "before I knew it - I had nothing."
During cross-examination, Mr Tron denied being a "very skilful and adept liar", but said: "I can lie when I need to."
During a National Crime Agency undercover operation, an officer was placed on the ferry and recorded conversations.
Prosecuting barrister Paul Mitchell accused Mr Tron of telling the truth to the undercover officer and "lying today because it's the only way you can save your skin".
Mr Tron claimed he had suspicions about the undercover police officer "within a few days", but played along by making up stories.
The trial continues.


More than 100,000 people have signed a petition against the closure of a beach on the French Riviera to allow Saudi King Salman to holiday in private.

The petition insists the public beach in Vallauris should be "available for the benefit of all".
Authorities sealed off the beach early on Saturday to prevent any protesters from occupying it as King Salman was due to arrive in the area.
The Saudi monarch is expected to stay at his villa for three weeks.
He and his entourage of about 1,000 people arrived at Nice airport on Saturday on board two Saudi Arabian Airlines Boeing 747s.
The king's inner circle is staying at the villa - between Antibes and Marseille - while about 700 others will stay at hotels in Cannes.
Michel Chevillon, president of an association of Cannes' hotel managers, said the visit was "clearly good news" for hotels and the local economy.
"These are people with great purchasing power," he said.
However, the closure of a section of La Mirandole beach beneath the king's villa has outraged many local residents.
"We recall that this natural zone, like all maritime public estates, is an intrinsic public property that should be available for the benefit of all, residents, tourists, French, foreigners or people passing through," the petition says.
"We ask the state to guarantee the fundamental principle of the equality of all citizens before the law."
The mayor of Vallauris has also written to President Francois Hollande in protest at unauthorised work carried out at the property.
A cement platform has been poured on to the beach to provide a lift up to the king's villa, although the Saudis have promised to remove it when they leave.
"We understand the security reasons and the nation's greater interest. But nobody can exonerate himself from the laws of the land," Mayor Michelle Salucki wrote.
There is also a ban on coming within 300m of the villa by sea.
"We're sick and tired of this messing around," one local woman said, quoted by AFP news agency.
"I can see it's normal that you need to guarantee their security, but they should let us go for a swim."


Former Respect Party leader Salma Yaqoob has announced she is standing in Bradford West as a candidate in the upcoming general election.

Ms Yaqoob will stand as an independent candidate in the constituency currently held by Labour.
She quit Respect in September 2012 blaming a "breakdown in trust" at a senior level with others in her party.
Bradford West was held by the Respect MP George Galloway until May 2015 when he was beaten by Labour's Naz Shah.
Ms Yaqoob stood in the 2010 General Election, where she came second in the Birmingham constituency of Hall Green. She stood down as a local councillor in 2011 after five years, citing health reasons.
Other candidates standing in Bradford West are:
In 2015, Naz Shah beat George Galloway into second place, with 19,977 votes compared to 8,557.


Former world champions Ben and Tom Birchall stormed to their fifth TT win in the opening sidecar race of the week on the Isle of Man.

The brothers from Mansfield set a lap record on their second lap of the race of 117.119mph, finishing 25.669 seconds ahead of John Holden and Lee Cain.
Dave Molyneux and Daniel Sayle finished third a further 18.717 seconds behind.
Ben Birchall said: "Good bike, great team, it's all there. It could not have gone better."
He added: "We could have gone quicker on the last lap but it got really wet at the top of the mountain.
"Thanks to everyone that helped us. Another win at the Isle of Man and it was perfect."
The Birchalls led Holden and Cain by 12.124 seconds after lap one, who in turn were 7.738 seconds ahead of Molyneux and Sayle.
They built their advantage to 24.934 seconds on lap two of the three-lap race - setting the fastest-ever lap of the Mountain Course for a sidecar.
Holden said: "Dave and I had a proper scrap out there and Lee was a proper star. It was a great race."
Molyneux completed his 50th TT, achieving his 30th podium.
Alan Founds and Jake Lowther finished fourth, Tim Reeves and Mark Wilkes fifth and Peter Founds and Jevan Walmsley sixth.
Subsequent practice sessions were cancelled on Monday because of rain and fog around the course.


Luton Town manager Nathan Jones says his side's remaining eight games must "mean more" to his players if they are to win the "shootout" for promotion.

Jones said the Hatters "don't know how to grind out a win" after their 1-1 draw with lowly Newport County.
Luton are currently in fifth place in League Two, five points off the automatic promotion places.
Jones said his team were playing too many "powder-puff crosses or little passes when we want a killer instinct".
He told BBC Three Counties Radio: "We've only lost once in four games, but we've only taken three points. Now we have eight games to go - it's a shootout."
Danny Hylton gave Town the lead after five minutes at Rodney Parade, but Sean Rigg levelled for the hosts, who remain second bottom of League Two.
Jones, who signed a new three-and-a-half year contract with the Hatters on Monday, added: "It has to mean more to the [the players]. It's livelihoods. It's promotion. It can make or break them."


A care home which was put into special measures in January is still breaching regulations, inspectors have said.

The government's watchdog the Care Quality Commission (CQC) found that Sandridge House in Ascot was still "inadequate".
Inspectors said that there were not "safe staffing levels" at the specialist dementia care home.
Care home bosses have said that a "national shortage of trained staff" has made recruiting tougher.
The 33-resident care home was inspected in December 2015 and was put into special measures.
After the most recent inspection, the CQC reported that it had "found a number of breaches" which impacted on patient safety.
Not enough had been done to address the extreme risks of people falling, the watchdog added.
Its report said: "People continued to have injuries resulting from falls. We observed this occurred due to failure to assess, mitigate and review risks for people at high risk of falling."
In December 2015, the Health and Safety Executive said that there was evidence of legionella in the water pipework.
The bacteria can cause a pnuemonia-like illness called legionnaires' disease.
Inspectors also said that some people at high risk of malnutrition or dehydration were not appropriately monitored.
Group manager Naila Nanji said: "The standard of cleanliness has improved considerably and there is particular attention to good practices and stricter guidelines within infection control.
"There is a national shortage of trained staff, caused in large part by government policies restricting work permits for trained nurses outside of the EU, which ultimately causes an adverse effect and an acute shortage."