Manager Tony Pulis has left Crystal Palace by mutual consent less than 48 hours before the start of the season.

The 56-year-old met chairman Steve Parish on Thursday as tension between the pair came to a head.
The former Stoke boss, who won the Premier League's manager of the year award last season, had been frustrated by the failure to land a number of summer transfer targets.
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Assistant manager Keith Millen will be in charge against Arsenal on Saturday.
On Friday, Palace issued a statement, saying: "The club would like to thank Tony for his efforts with the club last season and wish him all the best for the future."
Bookmakers' odds had tumbled on Thursday as speculation over the Welshman's job intensified.
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Malky Mackay and David Moyes were suggested as potential candidates to replace Pulis by former Palace defender Matt Lawrence, speaking on BBC Radio 5 Live.
The remainder of Pulis's coaching staff - including Gerry Francis and Dave Kemp - have been asked by Parish to stay on for the trip to Arsenal before leaving the club.
Gunners boss Arsene Wenger said:  "He has done a remarkable job last season, made a miracle. I was at a managers' meeting on Monday, he was there and nothing indicated he wouldn't be here tomorrow."
Pulis wanted to assume greater control of all football matters at Selhurst Park, especially transfers, after missing out on a number of key targets.
He and Parish also disagreed about their vision for the club.
Parish and sporting director Iain Moody are in charge of delivering the potential signings earmarked by Pulis.
Moody arrived in November from Cardiff City, where he had held a similar role before being dismissed by the owner Vincent Tan.
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Palace signed Liverpool full-back Martin Kelly on Thursday, while striker Fraizer Campbell arrived from Cardiff for £900,000 and former Fulham centre-back Brede Hangeland have joined on a free.
Just hours before Pulis left, Kelly told the club's official website: "I'm looking forward to playing under Tony and playing regular football again."
Hangeland also picked out Pulis as one of his reasons for moving to Selhurst Park, saying: "All the things I've heard about this club - the manager, players, staff - it's all been good."
Pulis left Stoke last year after seven years at the Britannia Stadium, having guided the Potteries outfit to the Premier League and kept them there, extending his record of never being relegated.
Palace were bottom of the table when he was named as Ian Holloway's replacement in November 2013.
Pulis managed to halt the decline, and a run of five successive victories from 29 March, starting with a 1-0 win over Chelsea, steered the Eagles to safety.
They famously came back from three goals down in the final 11 minutes against Liverpool to all but deny the Reds the title at the end of last season.
Their eventual 11th-placed finish was the club's highest since 1992.
Former Palace chairman Simon Jordan told Talksport that managers "have to do as they're told".
"Football managers are incredibly difficult people to deal with because they believe their own hype," Jordan said.
"I know what managers will do, they will put you to the sword. If you don't stand up and you don't manage your football club properly, they will run havoc in it and then they will disappear into the ether when it least suits you."
Pulis's transfer spend since joining Palace is approximately £10.5m - including January signings Scott Dann, Jason Puncheon and Wayne Hennessey, who arrived in a £3m deal from Wolves, and loan deals for Tom Ince and Joe Ledley.
At the end of last season, Parish spoke to the South London Press about Pulis, saying: "He took a big chance coming to Crystal Palace on a verbal promise that I and the club would support him financially."
After steering Palace to safety, Pulis said: "In the first couple of weeks Steve and I had our moments but the longer the season has gone on the closer we've got.
"Our relationship, which I think is the most important at a football club, has got more and more solid."
Hull defender Curtis Davies tweeted: "Can't believe Pulis has walked. Done a great job in his short time at Palace. Who do you think will replace him? Malky Mackay a good shout?"


More than 200,000 cups made from stainless steel smelted in Sheffield will be handed out to revellers at Glastonbury Festival.

The cups are being tried out as part of organisers' efforts to reduce the amount of waste produced on the site.
Festival-goers will pay a deposit for one of the cups to use throughout the event before returning, donating or keeping it at the end of the weekend.
The cups were developed by Sheffield-based Glogg and made in Birmingham.
Festival organiser Michael Eavis: "For me, the single most important thing was being able to source British stainless steel for the cups from the place where it was invented - Sheffield, and then to take it on to the home of manufacturing - Birmingham.
"Week after week, there's a story in the national press about jobs in the UK steel industry being put at risk.
"There's seemingly no end to the negative slide of this critical industry, and if the jobs, skills and infrastructure are lost they won't be replaced.
"We've worked on this project over the last three years, which will hopefully encourage other UK businesses to think about how they can support our steel industry during these very challenging times."
The cups will be in use at 10 bars across the site, with customers paying a Â£5 deposit when they buy their first pint.
The festival's green initiatives and sustainability co-ordinator Lucy Smith said that at the end of the festival the deposit would either be refunded or donated to charity, depending on whether a cup is returned or kept.
However, she said she hoped that those revellers who chose to keep their cups would not let them gather dust.
She said: "Don't put it on a shelf and never use it again: the idea is that when they go to another event they take it with them and reuse it."


A man has appeared in court charged with murdering a man at a flat in Brighton on Christmas Day.

Oliver Parsons, 30, of no fixed address, appeared at Guildford Crown Court accused of killing Joe Lewis, 24, of Rustington, West Sussex.
Mr Lewis was stabbed at a block of flats in Montague Street at about 01:00 GMT and died at the scene.
Mr Parsons was remanded in custody to appear at Lewes Crown Court on 20 March.
He appeared by video link from Lewes Prison.


Huawei has announced its first Windows 10 tablet, at Mobile World Congress in Barcelona.

The MateBook is designed to be an alternative to laptops and is thinner than Microsoft's rival Surface Pro 4 thanks to its use of a USB-C port.
The Chinese firm is already the world's third-bestselling mobile phone-maker, but until now had only made tablets powered by Android.
Experts said it made good business sense to expand into the new category.
However, there was no mention of the much-rumoured P9 smartphone at Huawei's event.
When questioned about the no-show, the firm told the BBC it did not discuss future products.
The MateBook features a 12in (30.5cm) display and can be clipped into an add-on keyboard and used with a stylus.
Its advantages over Microsoft's Surface Pro 4 are that it is:
However, the disadvantages are that Huawei's machine is only offered with a sixth generation Core M processor, and not Intel's more powerful Core i5 or i7 chips.
In addition, the decision to rely on a USB-C port means that older peripherals cannot be attached directly to the machine and it lacks a built-in display port.
In order to gain USB 2 and external display connections, users will need to carry an adapter, which also includes an ethernet port. The tablet does, however, have a built-in SD memory card slot.
One other unusual feature about the machine is that its stylus includes a laser pointer at the opposite end to its nib.
One company watcher said he was "totally underwhelmed" by the product, but said he could understand why it had been developed.
"I expect that Huawei will receive contributions to its marketing costs from Intel and Microsoft," said Ben Wood from the CCS Insight consultancy.
"Both of them will want to go to great lengths to help it promote the product, and that takes out some of the risk for Huawei.
"I should add that it has also been beautifully engineered. But even so, this is a segment of the market that is getting very crowded very quickly with Lenovo and Samsung also having recently announced similar products."
Apple also put its iPad Pro on sale in November, which likewise has a stylus and clip-on keyboard but still uses the firm's mobile iOS operating system.
PC shipments are forecast to fall further this year - research firm Gartner has predicted they will decrease by about 1% - and tablet shipments as a whole are expected to see an even steeper drop.
But the so-called "detachable tablet" category, which includes the MateBook, is growing.
Sales were up 75% in 2015 and are forecast to double again this year, according to market intelligence firm IDC.
"It's a product category that we are going to see more people get involved in as it's a good way for the mobile players to attack the PC market," commented Tim Coulling, an analyst at the tech consultancy Canalys.
"But the thing is, Microsoft Surface Pro benefits from the company's brand name and also  makes use of the firm's business-to-business sales channels."
These are things where Huawei's MateBook could be at a disadvantage, he added.
The entry-level MateBook will cost $699 (Â£485) when it goes on sale later this year, making it $100 cheaper than the basic Surface Pro 4.
To purchase its keyboard, MatePen and MateDock will cost an additional $277.
Follow all the BBC's MWC coverage via the "Mobile World Congress 2016" tag in its news app.
You can also keep track of our technology correspondent Rory Cellan-Jones and the rest of the team covering the Barcelona event via a dedicated Twitter list.


A railway guard has been cleared of endangering passenger safety after an elderly woman fell between a train and a platform.

Edna Atherton was 88 when she fell from the platform, suffering broken ribs and a cut head at Birkenhead's Hamilton Square station in July 2015.
The prosecution had claimed Martin Zee, 33, had not followed Merseyrail's door closure safety procedure fully.
But the defence argued the fall was an accident, Liverpool Crown Court heard.
They said there was a "fundamental failing" in the 17-point safety procedure itself, because the curve of the platform meant there were several blind spots.
The trial was shown CCTV footage of Mr Zee leaving the back of the train and walking to the wall of the platform to check all passengers had got off and on the carriages.
After Mr Zee pushed the button to close the doors, two women can be seen trying to board the front of the train. As the doors were reopened and closed, one of them fell between the train and the platform.
Mr Zee, of Coronation Road, Runcorn, Cheshire, had pleaded not guilty to a charge of endangering the safety of railway passengers by wilful omission or neglect.
The jury cleared him after two hours of deliberations.
A Merseyrail spokesman said its internal investigation and another by safety regulator the Office of Rail and Road "found there was no case to answer".
"The door closure warning alarm is in place to alert passengers to the danger of closing doors and we rely on passengers heeding that warning and not attempting to board or alight while the doors are closing."
A spokesman for British Transport Police said: "It is our duty as a police force to investigate and present the evidence to the Crown Prosecution Service."
A Crown Prosecution Service spokesman said: "The decision to charge was made following detailed consideration of the evidence and in accordance with the Code for Crown Prosecutors."


Philip Pullman's novel The Firework-Maker's Daughter is to be turned into an opera for children.

Five singers and two puppeteers will tell the tale of Lila, a girl who wants to make fireworks like her father.
Pullman, a former teacher, originally wrote the story for a school play and published it in book form in 1995.
The opera will be directed by John Fulljames, associate director of opera at the Royal Opera House, and will receive its premiere in Hull in March.
It will then go on tour, taking in the Royal Opera House in London and the New Victory Theater in New York as well as Huddersfield, Watford, Bury St Edmunds, Buxton, Oxford and Newcastle.
The music will be written by award-winning composer David Bruce, working with librettist Glyn Maxwell.
Opera North's education director Rebecca Walsh said it "promises to be an explosive introduction to opera for families".
Frederic Wake-Walker, artistic director of The Opera Group, said: "The Firework-Maker's Daughter continues The Opera Group's commitment to giving people, and especially young people, their first experience of opera.
"It promises to be a wonderful show full of colour, humour and emotion."
The show will be co-produced by Opera North and The Opera Group in association with the Royal Opera House and Watford Palace Theatre.


BBC Sport tracks all the manager ins and outs as well as listing all the current bosses in the Premier League, Scottish Premiership, English Football League and National League.

All of the managerial movements for June will appear below, followed by the full list of each club, league-by-league.
To read the list for June, visit the ins and outs page.


Specialist anti-slavery teams are to be based inside UK airports in a bid to clamp down on human trafficking, ministers have announced.

The first team will be based at Heathrow from 1 April before the scheme is rolled out to other airports.
They will be tasked with identifying victims and disrupting criminal gangs involved in international trafficking.
The government says the scheme will ensure there is "no easy route into the UK for traffickers".
The new teams, which will be led by specially trained Border Agency officers, will also begin work at Gatwick and Manchester airports later this year.
The government has said "further ports" will be identified over the coming months.
Home Office minister Karen Bradley said Border Force officers could be the "first authority figure in the UK to have contact with a potential victim of modern slavery".
"Their role is vital in identifying and protecting victims and ensuring there is no easy route into the UK for traffickers", she said.
"The new specialist teams will build on existing skills and joint working and extend that expertise around the country."
The teams will be supported by the National Crime Agency, which will bring its child protection expertise in cases involving children.
The new system will also provide child victims with an independent specialist advocate with expertise in trafficking to act as a single point of contact.
Officers from the Border Force will receive specialist training to help identify potential victims.
The new scheme aims to ensure intelligence is shared and other bodies, such as social services, are kept informed of cases.
The announcement comes as the government aims to pass new legislation to fight slavery.
The draft Modern Slavery Bill could increase the maximum custodial sentence for human trafficking offenders from 14 years to life.


Residents who were evacuated from a nursing home in Fife after a major fire are being moved to another home in Glenrothes.

The elderly residents were initially cared for in hotels after the blaze broke out at Forth Bay Nursing Home in Kincardine on Monday afternoon.
The majority of residents and staff are now being accommodated by the council at Napier House in Glenrothes.
Fife Council said five people were taken to hospital with minor injuries.
About 60 firefighters used water jets to tackle the fire at the home in Walker Street, which broke out at about 15:20.
Members of the public came to the aid of the residents by helping them to find shelter.
Police Scotland said all residents and staff had been accounted for.
Part of the nursing home roof has been destroyed in the blaze.
Forth Bay is home to up to 53 residents, some of whom have dementia or complex health conditions.
Caring Homes, who run the nursing home, said all the residents were evacuated safely.
Michael Kellet, director of the Fife Health and Social Care Partnership, thanked all those involved in the recovery operation after the fire, including volunteers from Kincardine Community Association.
"Staff from services across the council worked until late in the evening to make sure everyone was taken care of properly and I'd like to thank them all for the dedication they have shown.
"Due to everyone's swift thinking, actions and planning we very quickly had arrangements in place to manage this challenging situation."
Mr Kellet added that planning was now under way to find longer-term accommodation for the residents.


Asad Rauf has been withdrawn from the Champions Trophy in June after media reports that the Pakistani umpire is being investigated by police in India.

"We feel that it is in Asad's best interests, as well as those of the sport and the event itself," explained International Cricket Council (ICC) chief executive David Richardson.
The ICC declined to comment on what the police are reportedly investigating.
Rauf, 57, has been on the ICC's elite panel of umpires since April 2006.
He was one of the umpires in the controversial Indian Premier League match that saw Rajasthan Royals players S Sreesanth, Ajit Chandila and Ankeet Chavan arrested for alleged spot-fixing.
A former first-class batsman, Rauf began officiating in 1998 and stood in his first one-day international in 2000.
He was elevated to the ODI panel in 2004, officiated in his first Test the following year and has now stood in 98 ODIs, 48 Test matches and 23 Twenty20 internationals.
The 2013 Champions Trophy will be held from 6 to 23 June in England and Wales, with matches held at The Oval, Edgbaston and the Swalec Stadium.


A fraudster destabilised the international art market by selling forged works of so-called "pitman painter" Norman Cornish, a court heard.

Richard Pearson, 56, from Sunderland, passed off 14 drawings and pictures to a gallery in Northumberland, leaving it more than Â£50,000 out of pocket.
He admitted fraud and forgery charges at Newcastle Crown Court and was jailed for three years and seven months.
Cornish was known for his paintings of industrial life in the North East.
The court was told Pearson was caught out after he made a "school boy error" when he used post-decimalisation prices on a fake receipt he claimed was from the 1960s.
Prosecutor Mark Giuliani said: "What was instantly and readily apparent was rather than being in pounds, shillings and pence it was in decimal pounds and pence."
The telephone number he used was also too long to be real.
Four of the fakes were sold on to private collectors, who the gallery in Corbridge has since had to refund.
Previously Pearson pleaded guilty to nine charges of fraud, two of forgery and two of using a false instrument with intent between December 2011 and February 2014.
Jailing him, Judge Edward Bindloss said the fakes were "convincing" and had caused confidence in the art market to diminish.
The family of Mr Cornish were present in court and in a pre-prepared statement said they hoped the conviction and the destruction of the fakes would restore confidence within the market.
Paul Currer, defending, said Pearson wanted to apologise for his behaviour and would pay back the money through a fleet of cars he gained from an inheritance.
Prosecutors said they would use the Proceeds of Crime Act to recover the money he owed.
Cornish, who died in August 2014, was a former miner who learned his craft at an art course run for pitmen at Spennymoor Settlement in County Durham.
His works have sold for five-figure sums.


Before there was Brexit - before there was a European Common Market or a Treaty of Rome - there was the sherry trade.

Shakespeare sang the praises of the golden fortified wine from the sun-griddled vineyards of southern Spain.
The English in particular have been drinking it by the barrel since Sir Francis Drake began looting supplies from the Spanish harbours he attacked.
The sherry wineries or bodegas in and around the southern city of Jerez de la Frontera are a handy vantage point from which to survey the trade landscape of post-Brexit Europe - an EU minus the UK.
On the one hand, times have been tough in recent years - sales of mass-market sherries have plummeted in the years since the 1980s, when every household in the UK kept a cheap bottle to hand, to pour into trifles on Sundays and relatives at Christmas.
On the other hand, the trade has historically showed extraordinary resilience at difficult historical moments, emerging successfully from World Wars One and Two, the Spanish Civil War and the Great Depression.
It was a point emphasised to me by Pedro Revuelta Gonzalez, in the elegant wood-panelled library of the Gonzalez-Byass bodega in Jerez.
He produced from the archive a ledger neatly recording the company's business in Britain during the early 1940s - the years of the Blitz and the Battle of Britain. The neatly-bound ledger, with handwritten entries in faded fountain-pen ink, was recovered from the wreckage of the company's London office, which was damaged in a German air raid.
Mr Gonzalez favours a soft Brexit - one which protects not only the interests of the sherry trade, but also those of the British expatriates who live on the coast, a short drive to the south, and the huge numbers of Britons who come every year to the resorts of Benidorm and Torremolinos.
"We believe in free trade," he told me. "And we hope and we're sure that the regulators in the UK and Europe also believe in that. So they'll try to find a solution that will be good for Britain, for Europe and of course for consumers, so there won't be big barriers in commerce."
If Brexit were to become a bitter affair, which involved British departure from the EU customs union and the single market, and strong restrictions on freedom of movement into the UK, then in theory at least those tourists in future might require a visa for their Spanish holidays.
Perhaps because this region of Spain has such close links with the UK, that confidence that a benign Brexit deal can be arranged seems fairly widespread.
What are the Brexit options?
Five models for post-Brexit UK trade
What does 'hard' or 'soft' Brexit mean?
How does leaving the EU affect expats?
In a party of British tourists on a tour of the Gonzalez-Byass bodega I met Mick Irwin and Glenys Nicholls, who told me they voted for Brexit. And if there are Brexiteers experiencing "Buyer's Remorse", as some commentators have suggested, then Mick and Glenys are not amongst them.
"No, no, no way - that's it, we've made our decision," Glenys told me. "I hope they don't go back and do another vote, because it's not the best of three until the other side get their way. We're coming out, come what way, and we'll just have to stand by our conviction and survive one way or the other."
When I asked Mick if there was some sort of contradiction between voting Brexit and then heading off to an EU member state on holiday he couldn't have been clearer.
"Not at all. They still want to sell us their sherry and they want to sell us their sun and sangria and they'll be cutting off their noses to spite their faces if they try to put British tourists off."
There was a general sense of surprise in Spain at the UK's vote to leave the EU.
Spain joined the EU only a decade or so after the death of the dictator Francisco Franco, and membership here is associated more with an anchoring of the democratic process and a modernising of the economy, than with unnecessary bureaucracy and untrammelled immigration.
Eugenio Camacho, the presenter and editor of a news programme on Radio Jerez, put it like this:
"The concerns about Brexit are mostly in terms of the historic commercial relations over sherry between Jerez and Great Britain. More than fear, there's uncertainty. At the moment when the consumption of sherry has started to recover in an important market like the UK this really puts the brakes on that recovery."
The process of negotiating the terms on which the UK leaves the EU has not yet begun. And it is possible that when it does it will be long and tortuous.
Southern Spain, with its strong historic British trading links, provides a reminder that when the EU is trying to settle on a negotiating strategy there will be voices calling from within for a soft Brexit and a soft landing - for what may be a difficult and protracted process.


After a mini-summit, European leaders have agreed steps to strengthen border controls along Greek and Slovenian borders, to try and slow the movement of refugees and migrants through the region.

There was grumbling, there was pleading, and there were several sharp retorts.
It was sometimes hard to remember that all the leaders who turned up here professed to want the same thing: an end to the chaos that increasingly marks the migration route through the Western Balkans.
They don't agree on the way to do it.
European Commission President Jean-Claude Juncker, who called the meeting, was hoping to rekindle a spirit of co-operation and trust that would allow countries to co-ordinate with each other - taking time to register new arrivals, and not simply waving them through to their next destination, or, as Slovenia has put it, "dumping" them on their border.
The pressure is real: 9,000 people a day have been crossing into Greece. And on Sunday alone, the government of Slovenia - a country of two million - said 15,000 people arrived at their border.
Without agreeing concrete steps soon, its Prime Minister Miro Cerar said Europe "will start to fall apart".
It's already splintering.
Deep rifts have opened up in the region, as the number of migrants has increased, and Mr Juncker's new proposals to strengthen Schengen's external borders in Greece and Slovenia play into national fears, as countries further along the route fear being turned into buffer zones.
In the story of Europe's migrant crisis, one nation's protection becomes another nation's problem.
In the end, he got a watered-down version of the co-operation he'd been hoping for: a commitment from countries to inform their neighbours before sending migrants and refugees their way, and to make an effort to process them properly.
In return, he promised extra border police for Greece and Slovenia, and 100,000 new places at reception centres along the West Balkans route.
Even before the meeting began, Croatia's Prime Minister Zoran Milanovic labelled the plan "unrealistic", drafted by someone "who had just woken up from a months-long sleep". The solution, he said, lay in Turkey and Greece.
It's a sentiment echoed by many of the leaders here - that without action from Turkey in stemming the number of people crossing its border into Greece, anything else is just tinkering around the edges.
The German Chancellor, Angela Merkel, acknowledged that the meeting was about managing the situation rather than tackling the root cause; any real solution, she said, would have to involve Ankara.
So was it all worth it? The Serbian Prime Minister, Aleksander Vucic, summed it up. One of the optimists going in to the meeting, he was the first to speak after it broke up.
Standing slack-limbed in front of the microphone, he told waiting reporters that the agreement would not be "immediately helpful".  We're looking at "small steps," he said.
"Hopefully, step by step, things will get a bit better."


Claudio Ranieri will get a special award from the Italian Football Federation for leading Leicester City to a shock Premier League title last season.

The 65-year-old Italian was sacked in February with the club in a relegation battle.
He will receive a Panchina d'Oro (Golden Bench) award next week.
Ranieri was named coach of the year at the BBC Sports Personality awards in December and also by Fifa in January.
The Italian FA, the FIGC, said: "During the ceremony a special Golden Bench will be given to Claudio Ranieri, the protagonist of the extraordinary journey by Leicester, with whom he won the Premier League."
The event takes place at Italy's national training centre Coverciano on Monday.


French police are investigating an assault on two Jewish brothers in a Paris suburb, in which the attackers allegedly shouted anti-Semitic abuse.

The eldest, 29, and his brother, 17, say one of the attackers was armed with a hacksaw and they both suffered cuts during the struggle.
They say they were harassed while driving through Bondy on Tuesday evening, wearing kippas.
Bondy is near Aulnay-sous-Bois, an area hit by anti-police rioting this month.
No suspects have been detained yet in connection with Tuesday's assault.
Bondy is part of Seine-Saint-Denis district, north-eastern Paris, where violence erupted earlier this month because of the alleged rape of a young black man by police.
The man, called Theo, left hospital this week after emergency treatment. Violence also rocked some other suburbs.
Seine-Saint-Denis has a high density of immigrants, many of them Muslims. The district suffers from high unemployment and urban deprivation.
A French group combating anti-Semitism, called BNVCA, said the two Jews had been racially abused first by two assailants who had followed them in a van.
In a statement to police, quoted by French commercial broadcaster LCI (in French), the younger brother says that when the van cut off their car, his brother lowered his window to ask why.
"I do what I want on the road," the driver allegedly replied. "You dirty Jews! You pr***s! You're going to die!"
The younger man says the driver then ran at his brother wielding a hacksaw, striking him with it as he tried to protect his face from attack.
When he saw the attacker aiming for his brother's neck, he struck him and a struggle ensued for the saw, in which his own fingers were cut, he said.
Some of the people then intervened to stop the attack, telling the brothers to leave, which they did, going to the casualty ward of a hospital, the younger brother said.
An eyewitness told LCI that he had taken a hacksaw off a man involved in the brawl but said he had seen no kippas.
"I saw neither Jew nor Muslim nor Christian nor atheist at the scene," he said. "As for the brothers' account, I heard nothing of the kind."
The BNVCA initially said that one of the brothers had lost a finger in the attack but they later corrected their account (in French) of the incident.
Anti-Semitism has been a feature of Islamist attacks in France in recent years:


Two more series of hit drama Line of Duty have been commissioned by BBC Two, one of several programmes announced to mark the channel's 50th anniversary.

The finale of the second series, starring Keeley Hawes and Vicky McClure, was watched by 4.1 million viewers last month.
Writer Jed Mercurio said he was "profoundly grateful" to fans for making the show such a "talking point".
Other commissions include a new series of James Corden's The Wrong Mans.
Both Corden and co-creator Mathew Baynton were nominated for TV Baftas earlier this week for their performances in the comedy thriller.
It is joined by a new comedy from Paul Whitehouse, based on the BBC Radio 4 series, Nurse.
It will star Esther Coles as a community psychiatric nurse, with Whitehouse playing many of her patients or their relatives.
Line of Duty - which revolves around a controversial police anti-corruption unit - will return for two more series of six hour-long episodes.
"For series three and four, I can promise two explosive new cases for AC-12, new guest stars as police officers investigated for corruption, further twists and turns from the loose ends of series two, and maybe even some surprise reappearances," said creator Mercurio.
Controller of BBC Drama, Ben Stephenson, said commissioning two more series of the "unique, powerful and gripping drama" was "the easiest decision I have made".
A documentary series on foster carers, War In Afghanistan and natural history series Beyond Human have also been announced for BBC Two.
The latter will explore how animals sense their world beyond human capabilities, focusing on a different sense in each episode.
Meanwhile, The World Made By Women will explore 20,000 years of women's history - from Empress Wu Zeitan to Margaret Thatcher - while several other documentaries will also look at the role of women throughout history.
They include a series on the suffragettes and a look at the Women's Institute, entitled The Real Calender Girls.
Women also feature heavily in Radio 2's latest line-up of special presenters for the spring, with Carla Bruni, Scissor Sisters star Ana Matronic and Sheila Hancock all fronting their own shows.
Carla Bruni's Postcards From Paris will see the model turned singer-songwriter and former first lady of France, presenting a three-part series taking listeners on a tour of the city and her favourite French music.
Ana Matronic delves into her disco record collection for a four-part series while stage and screen star Hancock will explore Music with a Message for two programmes.
"I'll be playing music which I hope will make listeners think, that can possibly change their thinking - it's music with ideas, music with a message," said Hancock.
The season will also feature a two-hour special with country queen Dolly Parton.


The family of former NHL player Steve Montador, who died earlier this year, are suing the league over accusations it failed to inform him about the risk of long-term brain damage.

The Canadian, who played 10 seasons with six teams, was found dead in his home in Toronto in February aged 35.
According to court papers, he had 15 documented concussions in his career.
The suit was filed on behalf of his son, Morrison, and other family members by his father, Paul.
Montador Sr said his son suffered from depression, memory problems and erratic behaviour and was involved in 69 on-ice fist fights during his NHL career.
"During regular-season NHL games, pre-season NHL games, NHL practices and morning skates prior to NHL games, Steven Montador sustained thousands of sub-concussive brain traumas and multiple concussions, many of which were undiagnosed and/or undocumented," claimed the suit.
According to Associated Press, several years before his death, the player had decided to donate his brain to the Canadian Sports Concussion Project.
In May, three months after his death, researchers confirmed Montador's brain had chronic traumatic encephalopathy (CTE), the brain disease that has been linked to repeated head trauma.
"The NHL continues to ignore the lasting problems caused by multiple head traumas suffered by its players," Paul Montador said in a statement.
"Tragedies like that of my son Steven will continue until the problem is addressed. The NHL knows, but denies, that years of repeated head injuries cause long-term brain problems."
NHL commissioner Gary Bettman declined to comment on the case, and said the league was not going to litigate the case publicly. Deputy commissioner Bill Daly said in an email the "claims made in the lawsuit are without factual or legal merit".
In April, the National Football League reached a settlement of a lawsuit brought by former players over concussions that could cost the NFL $1bn (£665m) and a US federal appeals court is considering whether the settlement figure is sufficient.
In Britain, rising fears over head injuries across all sports prompted the Football Association to issue new guidelines earlier this month to all involved with the game on how to deal with concussions.
World Rugby's chief medical officer Martin Raftery recently told the BBC the sport's rules may have to change to reduce concussions. with reported concussions doubling in five years.


BBC 6 Music are getting behind the Make Your Move campaign and want you to join them.

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Early breakfast presenter, Chris Hawkins, and weekend morning DJ, Mary Anne Hobbs, have picked up tips from Make Your Move personal trainer, Martin Cullen, and are encouraging listeners to get a little more healthy. See the week in workouts below.
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This is a great one for your posture as well as toning up your whole arms.
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This move particularly works the backs of the arms, also know as the triceps or 'bingo wings'.
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This move will focus on toning your upper arms.
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This move will give you definition in your triceps as well as working your back muscles
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This intense and effective isolation exercise will help to develop powerful and defined triceps.
Follow Chris as he and Martin talk you through the moves, or read through the steps on Make Your Move.
From Monday to Friday this week (18-22 July), Chris is asking listeners to take on the Breakfast Time Toning challenge during the WorkOUT song at 05:40 BST each day.
This posture-fixing, bingo-wing-busting routine uses baked bean tins from the kitchen cupboard to get your upper body warmed up and ready for the day ahead.
Chris will have demonstrations of each day's routine for you right here  - just take a look below for your instructions.
You can play along by using the routine or you can do whatever exercise you want for the duration of the WorkOUT song just as long as you are moving!
Is it time to Make Your Move? Find out about our new campaign promoting accessible, physical challenges for everyone here.
Use #MakeYourMove on Twitter, Facebook or Instagram to show us your efforts. Videos or photos are welcome, we just want to see you make your move.
But there is no rest for the wicked (or for folk with good taste in radio).
Over the weekend, Mary Anne is encouraging her listeners to take a hike.
Wherever you are around the UK, Mary Anne wants to see where you are reaping the benefits of a good walk and will be asking for your most scenic pictures and videos from your morning walk.
Happy dog walking is always encouraged.
There will be a short film with everyone's contributions to the Make Your Move campaign, so you might just see yourself on iPlayer if you play along and use #MakeYourMove.
So, get up and get moving with BBC 6 Music this week!


The largest proportion of happy people in England are living in Mid Sussex, the smallest in Wolverhampton, according to official statistics.

Data collected over three years has been used to measure the wellbeing of more than 300,000 people across the UK.
On average, people rated their happiness between seven and eight out of 10.
But the largest percentage of people rating themselves at nine or 10 was in Mid Sussex (39.3%).
That compares with 21.8% in Wolverhampton.
Sara Erriadi, 27, from Bilston said: "I don't have any family here at all so for me personally to be surrounded by people I love would make me happier.
"But for everyone else - if we had a bit more sunshine or more things to do - like a fair - that might make people happier."
Joyce and Val from Tipton had travelled to Wolverhampton on the tram for a shopping trip.
Val said: "A place is what you make it. If you're a happy person you'll be in a happy place."
Joyce said: "I always find the people really friendly in Wolverhampton.
"There's happy people and there's miserable people wherever you go."
Councillor Garry Wall, leader of Mid Sussex District Council, said: "Being happy isn't just about where you are; it's also about who you are with and how you live your life. We live in an area surrounded by beautiful countryside, areas of outstanding natural beauty and enjoy a growing local economy but crucially we have strong local communities."
The ONS found ratings of life satisfaction and happiness were at their lowest, on average, for those aged 45 to 59.
Across the UK as a whole people in the Scottish Highlands came out happiest, with 43% saying they were a nine or a 10 on the day they were asked.
In Bolsover in Derbyshire, 17.2% rated their happiness between zero and four out of 10.
However, it still had far more people at the happiest end of the scale as well, with 37% giving themselves nine or 10 out of 10.
Liverpool and Wolverhampton were the only places where average happiness was below seven out of 10, at 6.96 and 6.99 respectively.
Despite its low happiness rating, however, Wolverhampton has also emerged as the place where people feel the least anxious, with 58.5% rating themselves at zero or one out of 10 for anxiety.
Mayor of Wolverhampton, Councillor Ian Brookfield, said: "On the one hand they suggest that people in Wolverhampton are less happy, yet by the same measure, there are fewer unhappy people here than in many other areas.
Research shows the better you fit into the personality of your area, the happier you are. Take the test to find the best place in Britain for you
Read more on happiness from BBC News:
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"At the same time, the report also shows that people in Wolverhampton have the lowest levels of anxiety anywhere in the UK, which appears to contradict the other findings. And of course, we need to remember this is a fairly small sample size of 1,690 residents over a three-year period.
"Wolverhampton and Black Country folk are, by nature, hard-working and self-deprecating - that's why in this area we are famous for our sense of humour - and this research shows that while we may sometimes be a little bit grumpy, we don't worry and get on with life."
Castle Point in Essex has the most people in England classing their anxiety at the higher end of the scale, with Derry and Strabane in Northern Ireland ranked the highest in the UK.
Cornwall has the most people in England who are satisfied with their lives. Asked to rate their overall life satisfaction out of 10, one third (33.3%) gave themselves a nine or 10.
The London Borough of Camden contains the lowest percentage of people who are fully satisfied with their lives, 16.7 per cent, followed by Wolverhampton on 16.9%.
The largest proportion of people rating their life satisfaction zero to four out of 10 is in Greenwich, at 9.22%.
Cornwall also scored the highest in England for how "worthwhile" people considered their lives, with 39.2% rating what they did either nine or 10 out of 10. However, it was sixth overall in the UK, behind the Scottish Highlands, Anglesey in Wales, the Scottish Borders, Armagh, Banbridge and Craigavon in Northern Ireland and Dumfries and Galloway in Scotland.
Harrow in London had the lowest percentage of people scoring themselves with the highest marks, 22.6%.


Sixteen-time champion Phil Taylor beat Vincent van der Voort to set up a PDC World Championship semi-final with Raymond van Barneveld.

Taylor, who last won the title in 2013, edged a tense see-saw match 5-3 despite losing three successive sets.
The 54-year-old said: "It felt more like survival than a victory but I'm through and that is all that matters."
Dutchman Van Barneveld trailed BDO world champion Stephen Bunting 3-2 but fought back to win 5-4.
The five-time champion broke Bunting's resistance with a 107 checkout to take a tense seventh set and followed it up with a 113 checkout against the darts at the start of the last set.
In the earlier contest, Taylor was far from his best, averaging 100.08 with just a 40% success rate with doubles, but was fortunate that Dutchman Van der Voort struggled with a back injury.
The turning point came in the sixth set when Taylor, trailing 3-2, won the decisive leg despite taking 8 darts to hit a double.


Albanian police say they have foiled an art-trafficking ring seeking to sell hundreds of works of art abroad.

Prime Minister Edi Rama said the artworks seized - reportedly more than 1,000 - could fill an entire museum.
Among them were paintings, icons and murals thought to have been taken from churches and cultural centres in south-east Albania and Macedonia.
Their value was estimated at hundreds of thousands of dollars, Albania's Top TV reported.
Welcoming the police operation, Mr Rama said the artwork had "risked joining the long list of works that have crossed the country's borders".
He said it was one of the largest operations against art trafficking, but that it marked just the beginning of a campaign in which he appealed to Albanians to "redress this lamentable plight of our heritage".
Experts say that Albania's Orthodox churches have been plundered of much of their art work since the fall of communism, and that the trafficking of stolen art is widespread.
"[Albanian society] has forgotten that this might be our temporary house, but it remains the perennial abode of generations to come and we owe it to them to pass on the country we inherited from our ancestors," the prime minister said.
State news agency ATA reported that two men aged 39 and 51 had been arrested, and that two other suspects were wanted.


A man who survived an attack by cows in which his brother was killed has told an inquest it appeared the herd wanted to "deliberately trample" them.

John Porter said he and his brother Mike, 66, from Edinburgh, had been walking dogs on leads when they were attacked in a field in Wiltshire.
He told Salisbury Coroner's Court they picked themselves up after being trampled only to be knocked over again.
There had been three previous cow attacks nearby, the inquest heard.
The fatal attack happened on a public footpath through Elbow Field in Turleigh in May 2013.
Mr Porter told the inquest that attempts to shoo the cows away failed and they were surrounded.
Mr Porter, from Monkton Combe, near Bath, said the cows seemed to "deliberately trample" on them "as if it was something they really wanted to do", adding that it was "extraordinary".
His brother, who lived in Edinburgh with Adrienne Sillar and their two sons, died at the scene.
The inquest heard there had been one previous attack in the field, in 2008, and two in a neighbouring field in 2011.
David Billington, from Winsley, described how he had been walking home one evening in October 2011 when he heard "thundering hooves". He was knocked unconscious and suffered a broken neck.
The farmer, Brian Godwin, 81, said the cows in the field were continental and British cross-breeds, which were "more excitable" than other cows.
He said he had put in place safety measures after previous incidents - including signs warning people to keep dogs on a lead and some electric fencing. Mr Godwin has since sold his cattle and is trying to establish a new herd, the court was told.
The inquest continues.


The Lake District has joined the likes of the Grand Canyon, the Taj Mahal and Machu Picchu by being awarded Unesco World Heritage status.

The national park was one of 33 sites around the world to be discussed by the Unesco committee in Krakow, Poland.
The committee praised the area's beauty, farming and the inspiration it had provided to artists and writers.
It is the 31st place in the UK and overseas territories to be put on the Unesco World Heritage List.
The committee suggested the impact of tourism be monitored and requested improvements in conservation efforts.
The delegates heard the 885 sq-mile (2,292 sq km) Lake District had been trying to obtain the Unesco status since 1986.
Lord Clark of Windermere, chairman of the Lake District National Park Partnership which put together the bid, described the decision as "momentous".
In pictures: Where are the UK's world heritage sites?
Men-only island gets World Heritage nod
"A great many people have come together to make this happen and we believe the decision will have long and lasting benefits for the spectacular Lake District landscape, the visitors we welcome every year and for the people who call the National Park their home," he added.
Steve Ratcliffe, director of sustainable development at the Lake District National Park, said the application had been a "long time in the making" and he was "incredibly proud" of the landscape which has been shaped by nature, farming and industry.
He told the committee: "The Lake District now becomes an international and global property and we look forward to working with you and our communities to make sure this site inspires future generations around the world."
About 18 million people visit the Lake District each year, spending a total of Â£1.2bn and providing about 18,000 jobs.
It is home to England's largest natural lake - Windermere - and highest mountain - Scafell Pike.
Nigel Wilkinson, managing director of Windermere Lake Cruises, said he was hopeful the Unesco status would put the Lakes on an international level.
"What we really hope is it will act as an economic driver and will grow the value, not the volume, of tourism by giving people more... reasons to make day visits and sustained visits."
Harriet Fraser, a writer and patron of Friends of the Lake District, said: "It's the most beautiful district but it has a very deep culture which is largely hill farming but also conservation."
Other UK Unesco sites include Stonehenge, Durham Castle and Cathedral, and the city of Bath.


The family of a Londonderry man who was left in a vegetative state after a violent sectarian attack have said they appreciate an apology by the PSNI chief constable.

Paul McCauley was attacked in 2006 by a mob at a barbecue in the Waterside. He has never regained consciousness.
His father, Jim, said the family felt the apology was an admission by police that there were failings in the case.
Only one person has ever been brought to justice.
Daryl Proctor, is serving a 12-year term for grievous bodily harm over the attack.
Addressing the McCauley family at a policing board meeting in the city on Wednesday, Chief Constable George Hamilton said:  "It's a tragic devastating story and I'm sorry that not more people have been brought to justice for the suffering that your son and your family have had to endure over the last eight years."
Mr McCauley told BBC Radio Foyle's News at One programme that he felt the apology showed that Mr Hamilton was "quite informed" about his son's case.
"He knew exactly what he was talking about and it also showed strength of the man," he said.
"He was quite happy to apologise for the failing within his organisation in the early stages of the investigation. We know the case has been passed to a higher grouping, with the serious crime squad now helping the case.
"The police are facing massive frustrations in getting the information that they deserve and that the  family deserve to get in order to put people behind bars."


A man accused of planting a fake bomb at a busy shopping centre has appeared in court.

The 21-year-old, of Portmead, Swansea, has been accused of placing an article with intent to cause alarm.
The alleged hoax saw Swansea's Quadrant Shopping Centre evacuated on 24 May - two days after 22 people were killed in the Manchester terror attack.
He was remanded in custody at Swansea Crown Court on Monday. A plea hearing is set to take place on 24 August.


Liverpool striker Danny Ings has been ruled out for the rest of the season with a serious knee injury.

Ings, 23, suffered a cruciate ligament injury in his first training session with new manager Jurgen Klopp, after making his England debut in Monday's Euro 2016 qualifier against Lithuania.
Ings has scored three goals for the Reds in seven games, since joining from Burnley this summer.
Liverpool full-back Joe Gomez is out for the season with the same injury.
The 18-year-old defender was hurt while playing for England Under-21s in their 3-0 win over Kazakhstan on Tuesday.
Ings scored Liverpool's goal in their most recent Premier League game, a 1-1 draw with Everton on 4 October.
He netted 43 times in 130 games for Burnley after joining from Bournemouth in 2011.
His Clarets debut was delayed for six months after he suffered a serious knee injury in August 2011, while a knee cartilage problem led to a two-month absence the following season.
Ings joined the Reds after his Burnley contract expired but because he was under 24, the Championship club are entitled to a compensation fee, which will be decided at a tribunal.
Liverpool, who travel to Tottenham in the Premier League on Saturday (12:45 BST), are already without striker Christian Benteke because of a hamstring injury.
The Reds are 10th in the table with only one win in their past six league games.


Gemma Arterton and Stephen Mangan have been announced as hosts of this year's Olivier Awards.

The ceremony, the biggest in the UK theatre calendar, takes place at the Royal Opera House in London on 13 April.
Olivier award-winner Julie Walters will provide the voiceover at the event.
Charlie and the Chocolate Factory and Stephen Sondheim's Merrily We Roll Along are the most nominated shows, with seven nods apiece.
Dame Judi Dench, Lesley Manville, Jude Law and Tom Hiddleston are among the acting nominees.
Arterton is to star in a West End musical version of 2010 film Made in Dagenham in November.
The former Bond girl will take the lead role of Rita O'Grady, a part played on the big screen by Sally Hawkins.
At the start of this year she played the lead in The Duchess of Malfi at the new Sam Wanamaker Playhouse in London.
Stephen Mangan's most recent stage outing has been in the Olivier-nominated Jeeves & Wooster comedy, Perfect Nonsense.
This year's Olivier set will be designed by one of this year's nominees, Es Devlin.
The stage designer was behind the London 2012 Olympic Closing Ceremony and, most recently, provided the set for the new musical I Can't Sing!
She is nominated for an Olivier Award this year for her designs for the play Chimerica.


The RSPCA investigated more than 400 allegations of animal cruelty per day last year - a rise of nearly 5% compared to the previous 12 months.

The charity looked into 149,604 complaints compared to 143,004 in 2015, but the number of successful prosecutions fell more than 6% to 744.
It said the surge in complaints may be due to increased use of social media.
Prosecutions included snakes decapitated with scissors and a golden eagle kept in a cramped kitchen.
Hotspots for complaints in 2016 were London, with 11,812, and West Yorkshire, with 7,920.
The regions subject to the most successful prosecutions were West Yorkshire, with 94, followed by North Yorkshire with 50.
Dermot Murphy, of the RSPCA, said: "I believe the figures show that we're not becoming crueller, but that people are simply less willing to stand by and do nothing if they think an animal is suffering.
"People are increasingly likely to share images or footage on social media of animals they believe are not being cared for properly."
Source: RSPCA. Figures relate to 2016
Source: RSPCA. Figures relate to 2016


The number of hostages killed or injured by Somali pirates increased significantly last year, according to a report by maritime officials.

The International Maritime Bureau says 35 people who were held hostage by Somali pirates died in 2011.
Levels of violence against seafarers in coastal waters around east Africa and the Gulf of Aden also remained high.
While the number of successful hijackings decreased, hostages are being held for longer, the report said.
Most of the hostages who died were killed while trying to escape during rescue operations.
The other deaths were directly at the hands of the pirates or due to malnutrition or disease during captivity.
While figures for hostages killed in previous years are patchy, the report says the 2011 figure does reflect a significant increase, and accounts for 3% of the total number of hostages taken.
BBC Africa analyst Mary Harper says it used to be rare for hostages to die in captivity since their value while alive has enabled pirates in the past to successfully obtain large ransom payments.
Source: 'Human cost of Somali piracy', International Maritime Bureau
The report also outlined some of the violence faced by seafarers and those taken hostage.
Last year nearly 4,000 seafarers were fired upon by Somali pirates, the report said
Of that number, 968 seafarers faced armed pirates who managed to board their vessels, while some 413 of those seafarers were rescued from secured rooms on their vessels by naval forces.
At least 1,206 hostages were held by Somali pirates in 2011, including 555 seafarers attacked and taken hostage during the year, and 645  captured in 2010 who remained in pirate hands.
Half of those held were subjected to punching and slapping and 10% suffered violent abuse such as being locked in freezers, burned with cigarettes and having their fingernails pulled out with pliers.
The average length of captivity also increased by 50% over the last year, up to an average length of more than eight months.
Some have been kept for more than two years, such as the crew of the Panama-flagged MV Iceberg who have been held hostage since March 2010.
The report says the ship's owner has gone out of business so there is nobody to negotiate the crew's release.


Prestatyn Town FC has been saved from closure after raising £53K to settle a winding-up petition from HM Revenue and Customs.

The former Welsh Premier League club - the unbeaten runaway leaders in the Cymru Alliance feeder league - had until Thursday to pay or face closure.
Chairman Chris Tipping broke the news to supporters on Wednesday night.
He was "unbelievably proud" of fans who donated more than 50% of the money after a four-week fundraising drive.
Prestatyn, who qualified to play in Europe after winning the 2013 Welsh Cup, have also raised cash from sponsored events, bucket collections, auctions and donations from local businesses.
The players have also contributed a "substantial amount" by donating their salaries.
The winding-up petition was issued in October following a two-and-a-half-year investigation by HMRC.
Mr Tipping said the club had been in a long-running dispute with HMRC about non-contract players' wages and expenses and that the amount owing included £17,000 worth of "unexplainable debt".
"This is not a case of us not paying our taxes. We are up to date with them," said Mr Tipping.
"This has been a dispute about HMRC refusing to accept that our non-contract players are not full-time employees.
"We got into a decent argument with them about wages and expenses and I've got three years' worth of paperwork to prove it. In the end they basically said 'we've had enough', gave us this bill and that was that."
The club was originally ordered to pay £75,000 for a period stretching back to 2011, but it was able to get that sum reduced as it was not recognised as a public limited company for the first two years quoted.
A spokesman for HMRC said: "We don't comment on the tax affairs of individuals.
"Our aim is to efficiently collect the debts due and to prevent things deteriorating further. We only initiate winding-up action where we believe this is the best way to protect both the interests of other taxpayers and creditors."
The 106-year-old club was relegated in 2015, ending a seven-year stint in the Welsh Premier League.
When Mr Tipping took over as chairman in May 2014, the club was operating at a substantial loss.
"Within three years, we've turned it around," he said. "At the end of this year we will pretty much break even."
"For us to have come so far and to be top of the league as well, I don't know how I'd have handled it if we had to close."
Mr Tipping admitted he was doubtful at first that the bill could be paid.
"I had to stand up in front of the fans at the first meeting in October and say 'look, this is achievable', but in your heart of hearts you think, 'this is almost impossible'.
"But one way or another, we've done it. It's really highlighted what the club means to so many people.
"A big thanks to everyone involved - this is probably our biggest result since winning the Welsh Cup."


A former Celtic FC youth coach has appeared in a court in Belfast charged with sexual activity with a child.

Jim McCafferty, 70, from Raby Street, south Belfast, is accused of offences alleged to have taken place between 2011 and 2014, when the alleged victim was between 13 and 16 years old.
Mr McCafferty is originally from Scotland.
A defence solicitor said he would not apply for bail because of concerns over the defendant's safety.
The judge remanded Mr McCafferty back into custody. He will appear in court again in January.
The defendant worked for several football clubs during the 1980s and 90s as a scout, kitman and youth coach.
The clubs included Celtic, Hibernian and Falkirk.
Mr McCafferty moved to Northern Ireland seven years ago.


Worcester Warriors have made Bath lock forward Will Spencer their third signing for the 2016-17 season.

Hampshire-born Bath Academy product Spencer, 23, follows the signings of centres Ben Te'o from Leinster and Jackson Willison from Grenoble.
"Will is a hungry young second rower who will add vital competition to the pack," said Warriors director of rugby Dean Ryan.
"We're looking to help him realise his potential here at Sixways."
Spencer will be reunited with Bath team-mate Tom Heathcote, who is in his first season as Warriors' first-choice fly-half.
"From speaking to Dean and Tom, Worcester feels like the right place for me to further my career over the next few seasons," said Spencer. "There is a great young squad at the club.
"Hopefully, over time, we can achieve our goals and become a top Premiership side."
Meanwhile, Worcester look like they could be without first-choice scrum-half Jonny Arr for the rest of the season.
Arr was an unexpected absentee for Saturday's last-gasp 32-22 defeat at Wasps. But it has now emerged that he has suffered a lateral media knee ligament injury - and Ryan predicts that he will be out for "three to four months".
After joining the Bath Academy in 2010, Spencer made the transition to Bath's first-team squad in 2014 and has since made 11 appearances.
The 6ft 7in forward spent a season on loan at London Welsh in 2013-14, playing a part in the Exiles' Championship-winning campaign.
His current contract with Worcester's Premiership rivals Bath was due to expire at the end of the 2015-16 season.


A seal that was captured after straying into a suburban Australian neighbourhood has been returned to the wild.

The fur seal made global headlines when it was photographed on top of a car in Launceston, Tasmania, on Monday.
The large male, estimated to weigh 200kg (441lb), was first seen in the middle of a street.
The animal was tranquilised at the scene and has since been driven to a river and released.
Wildlife biologist Rachael Alderman, from the Marine Conservation Program, said the relocation happened without incident after the sedative wore off.
"It ended well apart from the poor person with the car," she told the BBC on Wednesday.
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Dubbed "Mr Lou-seal" by police, the mammal is believed to have travelled to Launceston - about 50km (31 miles) from the sea - through nearby waterways.
Dr Alderman said fur seals were spotted regularly in Tasmania at this time of year, which is breeding season.
"There's changes to hormones, so we do get a spike in these large males turning up in places that are a bit unusual," she said.
"I guess the thing that was not standard (this time) was a that it was on a car bonnet and the amount of media interest that it generated."
William Gregory, who was staying at the Launceston home, said the seal caused minor damage to the car.
"We got up and there was this great big seal on the roof of the car which is definitely not what you'd expect on Boxing Day," he told the Australian Broadcasting Corporation.
"We'll replace the windscreen and pop a few dents out, they're just old cars so it doesn't really matter. It's worth it."


Glasgow's Scotland flanker Ryan Wilson has been cited for allegedly grabbing the testicles of Northampton's scrum-half Lee Dickson.

If found guilty of the incident, which occurred in the 25th minute of Sunday's European Champions Cup game, Wilson, 26, could face a minimum 12-week ban.
Wilson's disciplinary hearing will take place on Wednesday, with any ban likely to rule him out of the Six Nations.
Glasgow lock Tim Swinson has also been cited after a yellow card for punching.
Swinson was sin-binned in the 74th minute, but his offence could bring further punishment after being highlighted by match citing commissioner Jeff Mark.
Wilson has won 15 Scotland caps since his debut against Wales in 2013, and featured in four games at last year's Rugby World Cup.
He is likely to be included in head coach Vern Cotter's squad for the forthcoming Six Nations, which will be announced on Tuesday, pending any disciplinary proceedings.
Under World Rugby guidelines, the offence of 'testicle grabbing or twisting or squeezing' carries a suggested punishment of 12 weeks at the low end of the scale, to 18 weeks for a mid-range offence, and anything from 24 to 208 weeks (four years) at the top end of the scale.
Wilson missed last year's tournament after being suspended for three months without pay by Scottish Rugby following a conviction for assaulting a fellow rugby player on a night out.


A 57-year-old man has been arrested on suspicion of murdering a woman whose torso was found in a suitcase.

The man is understood to be Vitautas Jokubauskas, who rented a room in a shared house in Peterborough where the suitcase was found on Friday.
Cambridgeshire Police said it was no longer searching for the Lithuanian national.
Police were called to the house in Mayor's Walk after other tenants reported a "pungent smell".
They found the woman's torso in a suitcase, which did not contain her legs, arms or head.
Police said the murder suspect had been arrested in Clipston Walk, Peterborough, at about 19:00 BST on Sunday.
The woman's body has been removed from the house and a post-mortem examination is due to take place at Peterborough City Hospital on Monday.
A police spokesman said it had not been identified but officers were keen to trace a woman, believed to be Mr Jokubauskas's partner.
She was described as being about 40 years old, slim, with black hair and also Lithuanian, he said.


Team Sky rider Geraint Thomas says his hopes of winning Tirreno-Adriatico are "over" after a disastrous team time trial on the opening stage.

Team-mate Gianni Moscon crashed after his front wheel collapsed as Sky finished 18th, losing significant time to their main rivals in Italy.
BMC Racing won the 22.7km stage in 23 minutes 20 seconds, with Thomas' team one minute and 42 seconds behind.
"It was Murphy's law - what could go wrong did go wrong," said Thomas.
"Races like this are won in seconds - we'll try to be aggressive and make up for this, but I think general classification is out the window now."
The week-long World Tour stage race runs until 14 March.


The Bank of England has forced banks to find a further £11.4bn in the next 18 months to beef up their finances against the risk of bad loans.

Banks will have to set aside £5.7bn in the next six months in case future economic shocks mean some borrowers cannot keep up their repayments.
A further £5.7bn will have to be found by the end of next year.
The Bank's Financial Policy Committee (FPC) suggested lenders had become complacent about their lending.
"Lenders may be placing undue weight on the recent performance of loans in benign conditions," the FPC said.
The committee has also taken action to stop banks getting around key tests which are designed to stop them lending too much to consumers.
The FPC's assessment is that the risks facing the financial system remain at a normal level for now.
But there are "pockets of risk that warrant vigilance" it said, in the Bank's half-yearly Financial Stability Report.
Lenders, the committee said, are relying too heavily on borrowers keeping up payments as well as they have recently, and banks and other lenders have started lending to people with weaker credit records.
The FPC highlighted rapidly growing consumer borrowing via credit cards, personal loans and, notably, car finance.
Collectively known as consumer credit, these forms of borrowing have grown by more than 10% in the past year, far outstripping the growth of incomes.
While the amount of borrowing for consumer credit is just a seventh of the size of mortgage lending, the amount lenders have to write off because it is not likely to be repaid is ten times greater than for defaulting mortgage borrowers.
In a news conference, the Bank Governor Mark Carney explained that the Bank was worried about those households who are heavily in debt.
But their borrowing, he said, had not in fact increased the threat to the general resilience of banks.
"We are reinforcing some of the protection [for banks]," he explained, by telling banks to add to their financial cushions.
He declined to blame people for borrowing more, and said that most personal borrowing decisions were reasonable.
However, he advised: "Borrowers should consider adverse scenarios as well as positive scenarios."
The Bank is bringing forward by six months a so-called "stress test" in respect of consumer credit, whereby lenders have to test their ability to withstand losses on loans that go bad and are not repaid.
It is also blocking lenders from getting around affordability tests for lenders designed to stop them over-lending on mortgages.
Banks and building societies are currently allowed to lend a maximum of 15% of their mortgages to homebuyers who take especially large loans of more than four and a half times their income.
The lenders have to scrutinise the borrowers to ensure they could still afford their repayments if the Bank of England raised its official base rate by three percentage points.
But some lenders have been assuming they would not in fact pass on all of that increase in higher standard variable rates, thus allowing them to lend slightly more.
Mr Carney said these lenders were not "gaming the system" but instead appeared to have forgotten some of the lessons of the recent past.
Despite these concerns, Mr Carney stressed that the UK financial system was far stronger than at the time of the great banking crash in 2008-09.
He said that since then, UK households had reduced their levels of debt and that it was only in the past 18 months or so that personal lending and borrowing had accelerated again.
"The resilience of the UK financial system has strengthened since the financial crisis," Mr Carney said.


South Africa left-arm spinner Aaron Phangiso has been reported for a suspected illegal bowling action.

"Phangiso's action will now be scrutinised further in accordance with the CSA regulations," Cricket South Africa (CSA) said in a statement.
Phangiso, 32, who is in his country's squad for the World Twenty20 in India starting in March, will undergo testing by the International Cricket Council.
South Africa open their campaign on 18 March against England in Mumbai.
Phangiso, who was in South Africa's squad for their recent T20 series win over England, was reported after helping Lions into the final of South Africa's provincial 50-over competition, when he took 2-38 in eight overs.
Phangiso was disciplined by CSA after allegedly offending crew and passengers on a flight from India last October while in an intoxicated state.
He was also forced to apologise after being caught by television cameras imitating snorting cocaine while in his team's dugout during the second T20 international against England in Johannesburg on Sunday.


Labour has a problem with anti-Semitism.

Remember the councillor who thought Hitler was the greatest man in history?
Remember the candidate who suggested asking the so-called Islamic State why it wasn't attacking Israel?
Remember the student who said a large part of the Oxford University Labour Club had a "problem with Jews"?
Imagine the reaction, Jewish figures say, if these were stories about any other minority.
They appear with politically damaging regularity.
Now, though, it seems change will come.
Activists and MPs have told BBC Radio 4's Today programme they are trying to toughen the rules on anti-Semitism, Islamophobia and racism.
Even if they fail, I'm told a party review by Labour peer Lady Royall into allegations of anti-Semitism may well also suggest rewriting the rulebook.
That is highly unlikely to be the end of this story, though, because it goes to the heart of a poisonous atmosphere in parts of the Labour movement.
Jeremy Corbyn's critics - who are many and vociferous - put the blame at his door.
It is he, they say, who has attracted new members with these views, who has condemned anti-Semitism but not acted and who has - on occasion - made what they think are utterly inadequate responses to offensive language.
Their concerns are fundamental - they simply don't believe anyone who describes the Palestinian militant group Hamas as a friend wants to tackle this issue.
On the other side, his supporters remind us of those condemnations of anti-Semitism and the decision to launch an inquiry, and point out this is not a new problem.
Lost in the fury about a member suspended then readmitted after anti-Semitic comments only to become vice-chair of her local party, was the fact those decisions were made not under Mr Corbyn, but his predecessor Ed Miliband.
No Labour leader could be expected to vet the thousands of new members who have signed up.
For some fighting Mr Corbyn's corner, this issue is serious and real, but is also being used as a stick to beat him by his internal political enemies.
There are those who have long seen allegations of anti-Semitism as attempts to silence legitimate criticism of Israel - on which different wings of the Labour movement take passionately opposing views.
The political debate, then, is not at heart merely a row about rules or party management, but fundamental differences within Labour.
Changing the rulebook will not necessarily change what people say or write, merely how they are dealt with.
It certainly won't change how people feel, and there are plenty in Labour who seek no part in its civil wars but wonder why anti-Semites choose to join their party.


David Cameron is to set out a string of new powers to tackle radicalisation, saying the UK has been a "passively tolerant society" for too long.

The PM will tell the National Security Council a counter-extremism bill will be in the Queen's Speech on 27 May.
The bill will include new immigration rules, powers to close down premises used by extremists and "extremism disruption orders".
Mr Cameron will say a "poisonous" extremist ideology must be confronted.
In other political news;
The proposals were first set out by Home Secretary Theresa May before the general election.
But the Conservatives were unable to secure the backing of their then Liberal Democrat coalition partners for the measures.
There is likely to be some opposition in the new Parliament on the grounds that some of the plans could infringe people's right to free speech, BBC home affairs correspondent Danny Shaw said.
The measures are also expected to introduce banning orders for extremist organisations who use hate speech in public places, but whose activities fall short of it being proscribed as a terror group.
Home Secretary Theresa May told BBC Radio 4 Today the government wants to "bring people together to ensure we are living together as one society".
She said: "What we are proposing is a bill which will have certain measures within it, measures such as introducing banning orders for groups and disruption orders for individuals, for those who are out there actively trying to promote this hatred and intolerance which can lead to division in our society and undermines our British values.
"But it will be part of a bigger picture , a strategy which will also have as a key part of it actually promoting our British values, our values of democracy, rule of law, tolerance and acceptance of different faiths."
The measures, she added, will focus on "extremism of all sorts... that is seeking to promote hatred, that is seeking to divide our society, that is seeking to undermine the very values that make us a great country to live in".
According to details given by Mrs May at last year's Conservative Party conference, such orders would apply if ministers "reasonably believe" a group intended to incite religious or racial hatred, to threaten democracy, or if there was a pressing need to protect the public from harm, either from a risk of violence, public disorder, harassment or other criminal acts.
The granting of a ban, which would be subject to immediate review by the High Court, would make membership or funding of the organisation concerned a criminal offence.
The extreme disruption orders could be imposed on individuals, using the same criteria.
Policymakers have debated the definition of extremism ever since Tony Blair's government looked at new laws after the 7/7 Tube and bus attacks in London a decade ago.
There are, potentially, two key challenges for the government in creating anti-extremism laws and tools.
First, can a definition of extremism that leads to someone facing restrictions, such as a ban on using social media, withstand legal challenges  - particularly on human rights grounds?
Secondly can such bans work in practical terms without tying up the resources of the security services.
MI5, for instance, already has triage-like systems to prioritise watching the most dangerous people: it can't monitor everyone with dangerous views.
That aside, this package of measures is part of a potentially significant shift in focus.
Ministers want tools to marginalise, restrict and silence these voices because disrupting their influence may buy time to intervene and bring someone back from the edge before it's too late.
Under the government's plans, the Charity Commission will be given more power to "root out charities who misappropriate funds towards extremism and terrorism", and broadcast regulator Ofcom will be able to take action against channels broadcasting extremist content.
The terror threat level was raised from substantial to severe last August in response to the conflict in Syria and Iraq.
Ministers responded by introducing new orders that can block British fighters from returning to the UK and give police the power to seize the passports of people suspected of plotting to join the fighting abroad.
Mrs May will tell the National Security Council - which meets weekly and is chaired by the prime minister - that the government will empower institutions to "challenge bigotry and ignorance".
Mr Cameron will say the new powers will make it harder for people to promote extremist views.
"For too long, we have been a passively tolerant society, saying to our citizens 'as long as you obey the law, we will leave you alone'," he will say.
"It's often meant we have stood neutral between different values. And that's helped foster a narrative of extremism and grievance."
The Conservative government will "conclusively turn the page on this failed approach," he will add, saying the UK must confront "head-on the poisonous Islamist extremist ideology".
Jonathan Russell from the Quilliam Foundation think tank, which challenges extremism, said the measures would tackle symptoms, not causes.
He told Today there was a danger of "negatively" altering the balance between national security and civil liberties.
And on the government's plans, he added: "I don't think it will tackle radicalisation. I don't think it will change the numbers of people who are attracted to this poisonous ideology. And I don't think it will attack the ideology itself."


Colombian media expressed surprise that voters rejected a peace agreement with the rebel Farc (Revolutionary Armed Forces of Colombia) rebel group on Sunday.

Newspaper headlines also reflected the uncertainty over the future of the Farc and the peace agreement.
Colombian daily El Tiempo wrote: "After the victory of the 'No', Santos calls for political dialogue and peace".
"The president summoned all political parties. Uribe is willing to cooperate in favour of a national pact," El Tiempo said.
El Espectador carried a simple but dramatic headline "Noooooooo", alongside a graphic showing how tight the result was.
"The rejection of the peace agreement almost kills the possibility of the Nobel peace prize," said El Espectador.
The newspaper also carried a story under the headline: "Colombians backing the 'Yes' vote heard the result of the plebiscite in tears".
El Pais asked: "And now what?"
The Cali-based paper also asked if a Nobel Peace Prize for President Juan Manuel Santos and Farc leader Timochenko was now out of the question: "Does the result kill off the chances of Nobel Peace prize for Colombia?"
El Pais also quoted analysts who the paper said "foresee a gloomy panorama for Colombia after the plebiscite".
"No to the accord," was El Colombiano's banner headline, with a picture of Colombian President Juan Manuel Santos and his sombre-faced cabinet reacting to the vote. It reported that the president was seeking a "national dialogue" to tackle the political crisis triggered by the rejection of the peace accord.
On its front page, El Heraldo wrote: "The Agreement with the Farc rejected by a divided Colombia".
The papers also carried a quote from President Juan Manuel Santos saying: "I will not give up; I will keep searching for peace until the last minute of my administration."
Other front pages voiced the deep uncertainty left by the vote. "And what comes now?" asked El Universal. "Now what?" wondered El Diario/La Tarde on its front page.
Despite the uncertainty and frustration reflected in the headlines, some outlets were more positive, focussing on the possibility that a national consensus could still be reached.
"Opportunity to seek national unity," was newspaper La Republica's headline. The business daily Portafolio took a similar view: "The option of a great national accord emerges."
Independent news website La Silla Vacia meanwhile concentrated on the power of the man behind the "no" campaign: "Uribe is still king".
RCN Noticias said on its website: "Santos: 'the bilateral and definitive cease fire continues in place", while Noticias Caracol ran a story quoting former Vice-President and "no" campaigner Francisco Santos saying: "The process should continue and the Farc should be given guarantees."
Caracol Radio's main story on its webpage said: "Santos and the Unity [National Unity party] are looking for reconciliation with Uribe supporters."
Colombian radio network W Radio took a different approach on its website by quoting Farc leader Timochenko saying "peace is still possible".
National radio network La FM said: "Colombia enters uncertainty after "no" wins the vote."
BBC Monitoring reports and analyses news from TV, radio, web and print media around the world. You can follow BBC Monitoring on Twitter and Facebook.


Lambeth Council has suspended negotiations over a planned Garden Bridge, saying not enough has been done to attract private-sector backing.

The proposed bridge would stretch from Temple to the South Bank in Lambeth.
But the council has said it will only allow planning to continue if it is assured that the project's funds are not taken from Transport for London.
The Garden Bridge Trust responded that private contributions will make up "the greater amount of funding".
London Mayor Boris Johnson approved plans for the Â£175m bridge in 2014 after Lambeth Council and Westminster City Council granted planning permission.
Westminster Council told BBC London the bridge was granted permission in light of its "iconic design and significant public benefits" and its position remains unchanged.
The chancellor and the mayor have pledged Â£30m each, with the rest due to come from private funding.
Lambeth Council asked for assurances that this money would not be taken from Transport for London.
The leader of Lambeth Council, Lib Peck, said: "During a time of severe spending cuts, putting Â£30m of public funding towards this new bridge isn't justified."
She claimed that since June there had been "little progress in attracting alternative funding from the private sector".
She went on to state that future mayoral support for the scheme is "weakening", a reference to Labour mayoral candidate Sadiq Khan's announcement that he does not support the project.
Mr Khan described the bridge as a "white elephant" which "no longer represents value for money".
Bee Emmott, executive director of the Garden Bridge Trust, said: "We are surprised at the letter, particularly given how closely we have worked with Lib Peck and Lambeth Council for the past two years.
"We have been very clear from the outset that the public funding is a way to stimulate private contributions, which will make up the greater amount of the funding."
She pointed out that a "significant" amount has already been spent in the preconstruction and said the Trust will seek clarification from Lambeth about its announcement.
A Populus poll of 1,005 Londoners commissioned by Labour councillor Kevin Craig suggests 36% oppose it and 19% neither support or oppose.
This contrasts with a similar-sized survey carried out by ComRes in July 2015 on behalf of the Garden Bridge Trust, which indicated 78% support the bridge.
Actress Joanna Lumley is among those who support the crossing, designed by London 2012 cauldron creator Thomas Heatherwick.
It is estimated seven million trips per year will be made across the bridge.


Protests continue to spread across US cities against the killing of black men by police, following recent deaths in Minnesota and Louisiana.

Roads were blocked and missiles thrown in Minnesota, while armed New Black Panther Party members confronted police in Baton Rouge, Louisiana.
Dozens of arrests have been made but the rallies were mostly peaceful.
The situation is also tense in Dallas, where five officers were killed by a black man during a protest rally.
Security levels were raised at the police HQ after anonymous threats were received but an all clear was given after a search of a car park for a "suspicious person".
The protests against police killings were sparked by the deaths of Philando Castile in Minnesota and Alton Sterling in Louisiana.
In St Paul, Minnesota, overnight, fireworks, bottles and rocks were thrown at police as the demonstration closed a main interstate highway, causing gridlock.
This is the fifth night of protests in Baton Rouge where there is not just anger, there is rage.
Each night the gathering has intensified and police have responded with officers in riot gear. The two sides faced each other, often blocking the main highway outside police headquarters.
Occasionally the force would line up and rush into the crowd to make several arrests. Earlier, hundreds marched to the Louisiana State Capitol and stood on the steps with clenched fists raised up high - a salute to black power.
Read more from Baton Rouge and St Paul: US police protests near boiling point
Police said some officers had been hurt by fireworks and a number of arrests made. Demonstrators said police fired tear gas and rubber bullets.
In Baton Rouge, several hundred protesters gathered outside the police department shouting "No justice! No peace!" and also at the convenience store where Alton Sterling was shot.
There was also tension as armed members of the New Black Panther Party stood face to face with officers in riot gear. Louisiana law allows weapons to be carried openly.
Dozens of people were reported to have been arrested and at least two firearms confiscated. Among those held was DeRay Mckesson, who has become a national voice for the Black Lives Matter movement, and two journalists.
Baton Rouge protester Marie Flowers told Associated Press: "Black boys are being killed and this is just the culmination of what has been going on for decades."
Several hundred protesters blocked roads and bridges in San Francisco and Fresno in California, hundreds more protested peacefully in West Palm Beach, Fort Lauderdale and Miami in Florida.
In New York, at least 20 people were arrested as hundreds of protesters marched through Manhattan.
Protester Lorena Ambrosio told Reuters: "I'm feeling very haunted, very sad and just angry that black bodies just keep piling and piling up."
A "Weekend of Rage" called in Philadelphia saw a six-hour march by a multi-racial crowd. One protester told an officer: "Without the uniform and badge, you are just like us," AP reported.
Other protests took place in Nashville, Indianapolis and Washington DC.
Founded: 1989, Dallas. Current chairman is Hashim Nzinga
Membership: Unknown. Has claimed a few thousand although independent estimates suggest lower. Nevertheless, its members are often very active in protests
Manifesto: Has a 10-point platform that includes advocating the power to practise self-determination for a "black nation", the release of all black people in prison, an end to police brutality against black people, and a call for all black people to form a united front, arming themselves for self-defence
What others say: Both the Anti-Defamation League and the Southern Poverty Law Center call the NBPP racist and anti-Semitic, defining it as a hate group
Relation to original Black Panther Party: None. The Black Panthers, founded in 1966, were a militant, socialist organisation advocating black nationalism but also had community programmes. They largely disbanded in the mid-1970s. Co-founder Bobby Seale has called the NBPP's rhetoric "xenophobic" and "absurd"
The protests have continued despite an effort by President Barack Obama to soothe the tension.
On a trip to Europe, he said: "First of all, as painful as this week has been, I firmly believe that America is not as divided as some have suggested.
"When we start suggesting that somehow there's this enormous polarisation, and we're back to the situation in the 60s, that's just not true. You're not seeing riots, and you're not seeing police going after people who are protesting peacefully."
On Thursday, five white police officers were shot dead by a black man, Micah Johnson, during a protest rally against the police killings. Seven other officers and two civilians were injured.
Johnson, who was himself killed during the assault, supported black militant groups who encouraged violence against police.
Dallas police chief David Brown said Johnson had told a negotiator that he had wanted to kill white people, especially white police officers, because he was angry about the recent shootings of black men by police.
Johnson, 25, who officials say acted alone, was killed by remotely detonated explosives that were sent into a car park where he had taken refuge after the shootings.


The United Nations representative in Yemen has said that the Saudi-led coalition is bombing "effectively, trapped civilians".

Civilians in the northern city of Saada are struggling to flee Saudi-led coalition air strikes targeting Houthi rebels, reports and aid workers say.
The UN also warned that the indiscriminate bombing of populated areas is against international law.
Air strikes have killed at least 1,400, more than half civilians, the UN says.
The medical charity MSF, say that fuel shortages mean many people can only flee on foot.
The UN's humanitarian coordinator for Yemen, Johannes Van Der Klauuw, said he was "deeply concerned" by the impact of the latest air strikes on northern Yemen.
"Many civilians are effectively trapped in Saada as they are unable to access transport because of the fuel shortage," he added.
The Saudi-led coalition says it consider all of Saada a "military zone". On Friday it dropped leaflets warning residents to leave.
Mr Van Der Klauuw said that the decision to target the entire province "will put countless civilians at risk".
In a statement he added that many of Saada's residents were effectively trapped due to a lack of transport and that the leaflet warnings did not absolve the coalition of its responsibility not to target civilians.
Teresa Sancristoval, the emergency coordinator for Doctors without Borders (MSF) based at Saada's al-Gumhary Hospital, described intense bombing overnight, with reports of up to 140 strikes hitting the city in total.
She said her team had been treating seven pregnant women who were in labour, but five of them fled due to the intensity of the air strikes.
"The population that remains is very scared and worried... The market, storage facilities and government buildings have been destroyed and many civilians are suffering the consequences," she said.
Up to 130 strikes took place across the country on Saturday targeting rebel buildings, weapons stores and camps, a coalition spokesman said.
A spokesman for the Saudi-led coalition, Brig Gen Ahmad Assiri, said it was in response to Houthi rebels firing shells into Saudi Arabia earlier this week, killing 10 people.
Hundreds of families have already fled Saada province, security officials and Houthi rebels say.
However, the Saudis have accused the Houthi rebels of hiding troops and weapons in civilian areas to prevent them being targeted by airstrikes.
Gen Assiri said the Houthi rebels were preventing residents from leaving areas under fire.  A Houthi spokesman, Hamed al-Bokheiti, later denied this.
The UN has called on all parties not to locate "military personnel and assets in densely populated areas, thereby endangering civilians."
Saudi Arabia on Friday offered a five-day truce, which would start at 23:00 local time (20:00 GMT) on Tuesday.
But senior Houthi official Mohamed al-Bukhaiti told the BBC that the ceasefire had not been formally proposed and the Houthis would not respond until a plan was properly laid out.
Sunni Muslim Saudi Arabia has accused Shia rival Iran of arming the Houthis, a charge Iran and the Houthis deny.
The Saudis have been sheltering the internationally recognised Yemeni President Mansour Abdrabbuh Hadi, who fled there in March.