When 12-year-old Reuben de Maid from Cardiff sang on US talk variety programme The Ellen DeGeneres Show, it was not only his voice that made an impact .

The Welsh youngster was on the show ahead of his appearance on US talent programme Little Big Shots, but, unlike other male performers, he turned up for his debut in full face make-up.
Not surprising then that, after his performance, Ellen wanted to talk not only about his singing but also about his perfectly contoured face, which had caused a problem with other boys in the past.
"Ever since I was eight, I was so interested [in make-up]," he said. "And I would watch my phone for hours on end just learning from tutorials, like blogs, and seeing how to do it.
"Before, I used to get bullied a lot. I had friends but not a lot. In my drama group I used to get hit, punched and kicked.
"I tried to brush it off but brushing it off doesn't work, so I stood up to them and I told my mum," Reuben explained.
The show's host Ellen DeGeneres praised him for his uniqueness and courage and presented him with a laptop, make-up and lights so he could start his own make-up tutorial channel on YouTube.
"Good for you for not stopping being who you are, that's really tough," she said.
"The fact that just because you want to wear make-up, that somebody thinks they have to hit you or even call you names, it's not hurting anybody, it's what you want to do. And you look fantastic."
Reuben's mum Vicky Thomas told the BBC she was very proud of her son.
"When he started doing make-up, people used to say to me I can't believe you let him do that, but he loves it and he is so passionate about it.
"If I didn't allow him to be who he is, he wouldn't be happy."
Men are making huge gains in the make-up industry. Brands including CoverGirl and Anastasia Beverly Hills now have male models to showcase their products.
Earlier this year Manny Gutierrez was hired as Maybelline's first male face to represent their brand.
By Rozina Sini, BBC's UGC and Social News Team


A Red Cross employee has been killed and another injured in a gun attack by suspected Islamist militants in Mali, the aid organisation says.

Armed men opened fired on an aid vehicle as it travelled from Gao to Niamey in neighbouring Niger to pick up medical supplies.
The International Committee of the Red Cross (ICRC) said in a statement that it was "profoundly dismayed" by the killing.
Islamist militant group Mujao told AFP that it had carried out the attack.
The nationality of the ICRC worker who was killed has not yet been released.
"He was driving to collect much-needed medical equipment for a Gao hospital," said Yasmine Praz Dessimoz, head of operations for North and West Africa.
"His death is not only a tragedy for his family and for the ICRC, it will affect the life and well-being of tens of thousands of people," she added.
The injured worker was a member of the national Mali Red Cross and is in a stable condition in hospital.
The vehicle the pair were travelling in was clearly marked with the Red Cross emblem and was later burned.
"The ICRC is concerned about the rise in violence against humanitarian workers, which is preventing them from coming to the aid of individuals and communities in dire need," Ms Praz Dessimoz said.
A spokesman for Mujao, Abou Walid Sahraoui, was quoted by AFP as saying: "We have achieved what we wanted with this attack."
Mujao, which stands for Movement for Oneness and Jihad in West Africa, was formed in 2011 to spread jihad across West Africa and it believed to be a well-disciplined group.
Mali's desert north suffers frequent militant attacks despite a French-led operation to drive out Islamist fighters in 2013.
Earlier this month, a rocket attack on a UN base in Kidal killed a Chadian peacekeeper and two children.


Lawyers for Shafqat Hussain argue that there was never a credible investigation into his age at the time he was convicted of killing a child in 2004.

Despite appeals from human rights groups, he was executed in Pakistan on Tuesday.
His lawyers claim he was 14 when he was tortured into a confession and charged with involuntary manslaughter more than 10 years ago.
But the Pakistani authorities have said Shafqat Hussain's legal team failed to submit sufficient evidence to show he was a juvenile.
His brother, Manzoor, was at the jail to receive his body after the execution.
"What justice system is this?" Manzoor said. "He was just two and half years my junior - he was a young man."
Shafqat Hussain's uncle, Mohammad Altaf, who was also at the jail when he was executed, said his long imprisonment was inhumane.
"This is cruel... unjust. He was sentenced 12 years ago and all the appeals were rejected in one year. If he had been in prison for two more years he would have served a life imprisonment term already."
Shafqat Hussain met his family one last time before midnight, then was hanged shortly before dawn at the jail in Karachi.
Legal challenges and civil society campaigns saw his execution postponed four times this year.
But in the end, all the appeals and lobbying were ultimately turned down, including two letters from the provincial governments of Sindh and Pakistani-administered Kashmir which both asked for a stay on the execution.
Shafqat Hussein's case may have caused an international outcry to get him off death row but inside Pakistan it was cause for debate.
Many here support the death sentence and see it as justice being served in capital crimes.
Amnesty International described his hanging as "a sad day" for Pakistan. The sentiment was echoed by many activists on social media.
Others however, criticised NGOs for wanting to stop the hanging of someone they described as a child killer.
The authorities insist that Shafqat Hussain's case was properly handled.
"[His] case  was given due process," Ashtar Ausaf Ali, minister of state and a special assistant to the prime minister on law, told the BBC.
Mr Ali also added that for the last few months the government has revised the case very closely and looked at it a number of times.
The Pakistani government partially lifted a moratorium on capital punishment in the aftermath of an attack on a school in Peshawar last December, in which more than 150 pupils and teachers were killed by the Taliban.
The attack shocked the Pakistani people and put the political and military leadership in a very tough spot.
Since then the government has been trying to prove it's taken measures to combat militancy.
At the time Prime Minister Nawaz Sharif said the country was in an "extraordinary situation" that needed "extraordinary actions".
Later in the year the authorities lifted the moratorium on all capital punishment.
Pakistan is believed to have the largest number of death row inmates in the world. More than 8,000 prisoners are awaiting execution - many of whom have been in jail for over a decade.
So far about 200 people have been executed since December - only a small fraction of them were linked to terrorism.
Legal charities and human rights organisations have criticised the move saying the death penalty will not combat terrorism.
"The very logic of this is wrong," said Shahzad Akbar, a fellow at the legal charity Reprieve. "How can you deter a would-be suicide bomber by the death penalty?"
But Mr Ali insists that it has acted as a deterrent.
"The last eight months when the death penalty has been carried out was better than the last eight years (of the moratorium)," he said.
However, he didn't give any specific crime rates before or after the lifting of the moratorium.
"This is the law of the land," Mr Ali added. "It should be respected by all and adhered to."
But lawyers say the difficulty lies in Pakistan's flawed criminal justice system.
"There are inherent problems within the Pakistani justice system - like due process, proper legal representation and examining evidence," said Shahzad Akbar.
"What worries me is that there are many cases like Shafqat's that we don't have access to."
"People who have been languishing in jail for years and unless their cases are opened and looked at properly - the government should not hang them."


The chairman of the Airports Commission has rejected Gatwick's criticism of its report recommending expansion at Heathrow.

Gatwick claims Sir Howard Davies's decision not to back a second runway at the airport in West Sussex was flawed.
Sir Howard has written to the government insisting issues raised by Gatwick were carefully considered before the report was written.
The government is due to decide which airport should expand by Christmas.
The Airports Commission spent three years hearing submissions, including those from Gatwick and Heathrow, about how best to expand airport capacity in the South East.
Its report, on 1 July, said there should be a third runway at Heathrow.
Gatwick had argued strongly that it should be allowed to expand instead.
It has said the economic argument for expanding Heathrow is flawed and Sir Howard was using passenger numbers that were "10 years out".
Sir Howard forecasts Gatwick at 40 million passengers by 2024 but the airport says it will hit that number this year.
In a letter to Transport Secretary Patrick McLoughlin, Sir Howard says the Airports Commission believes strongly its work is robust.
"The majority of the points made by Gatwick Airport Ltd were also made to the commission in the course of its work and we considered them carefully before we reached our conclusions," he said.
Crispin Blunt MP for Reigate, who chairs a group of local MPs who oppose expansion at Gatwick hit out at the airport for continuing to campaign for expansion.
"Sir Howard Davies has demolished Gatwick's arguments, which amount to a feeble public relations effort to undermine the credibility of the Airports Commission's work and recommendations," he said.
Gatwick said it stood by its view that Sir Howard's report was flawed.
"If backed by government it will inevitably lead to 'Heathrow Groundhog Day' with nothing happening once again," said chief executive Stewart Wingate.


A Roman mosaic dating from the 4th Century in the first object to be installed in a new museum due to open in West Sussex in the summer.

The Chilgrove mosaic, discovered at Chilgrove Roman Villa, had to be moved in four sections, from Chichester District Museum to the Novium.
The museum has been built to display the remains of a Roman bath house.
Chichester residents can win the chance to visit the museum the day before it opens to the public on 8 July.
The Chilgrove mosaic will be sited opposite the remains of the bath house, which were excavated in the 1970s.
They were later preserved under a temporary car park but it was always the hope of the city council that a museum would one day stand on the site.
"The public will be able to see some of Chichester's extraordinary heritage in a wonderful setting," said Councillor Myles Cullen.
"The new museum is magnificent. It is sure to become one of the jewels in Chichester's crown.
"Not only will the museum attract local people but also visitors from far and wide. It will give a major boost to the local economy."
Novium will have about 150,000 artefacts, both on display and in storage.
"Together these tell the story of the district and the people that have lived here and influenced its development," said Novium manager Tracey Clark.
Residents will be able to apply for one of 40 "golden tickets" to preview the museum.


A desperate search is under way for an endangered African penguin "freed" by activists who may have unwittingly sent the bird to his death.

Buddy was stolen from a South African marine park by two students who claim it was a demonstration against animals being kept in captivity.
But Buddy was born in the park, and has no idea how to survive in the wild.
Experts say the penguin could last for just two more weeks before he will starve to death.
Dylan Bailey, manager of Bayworld in Port Elizabeth, said: "He is completely ill-equipped to survive in the wild. He will have no idea where he is.
"Luckily, he was a very healthy penguin - actually quite fat - so he has a good few weeks of reserves."
However, the park estimates that still means he has just three weeks from the moment he was stolen.
Buddy's disappearance was discovered the day after he was taken - when he could not be found for his monthly check up.
CCTV revealed two students - who have not been named, but are in their early 20s - climbing into the pool, and then bundling Buddy into the boot of a car before driving him a short distance to the sea, where they released him.
The two students had no intention of harming the bird, but their actions may have far reaching consequences, Mr Bailey told the AFP news agency.
Buddy was part of a breeding pair, and was looking after his two new chicks with mate Francis when he was stolen.
Since his disappearance, one of the chicks has died, although park officials do not know if this was related. Francis is also now unable to leave the nest, as Buddy is not there to take over from her.
Worse, African penguins are about to be added to the critically endangered list, with fewer than 20,000 breeding pairs in the wild. A successful match like Buddy and Francis is important for the species' very survival. Should Buddy not be found, pairing Francis with another mate could be close to impossible.
Mr Bailey explained: "They are monogamous animals. They pair for life. If we can't get Buddy back, we will try to pair her with another bird, but it may not be successful."
Staff have been scouring hundreds of kilometres of coastline, in the hope that one of them might spot Buddy.
"We are optimistic," Mr Bailey said. "We have hope he will come ashore before he becomes too weak."


There has been a small earthquake on the Lancashire coast, the British Geological Survey (BGS) has confirmed.

A small earth tremor was felt in the Blackpool area at about 0330 BST, it said.
Lancashire Police said calls had been received from several people who said their properties were shaking.
A BGS spokesman said the 2.2 magnitude tremor was thought to have centred on Poulton-le-Fylde. No damage or injuries have been reported.
A police spokesman said they confirmed reports from 10 callers when the force's control room on Bonny Street, Blackpool, began to shake.
Officers were sent to check reports of cracks on the road over a bridge in Lytham Road, but learnt they appeared up to two years  ago.
A spokeswoman for Blackpool Council said structural engineers would assess if there was any fresh damage to the road but it remained open to traffic.
She added there were no reports of any damage elsewhere in the borough, including at the council-owned Blackpool Tower.
Dave Thornton, a gift shop owner from Thornton Cleveleys, was woken up by the quake.
"All my wardrobe doors were rattling and I couldn't work out what it was.
"I went to the window, but everything seemed normal. It was a relief to be honest to learn that it was a quake because at least I know there's nothing wrong with my house," he said.
According to Brian Baptie, a BGS seismologist, the UK experiences about 20 earthquakes a year of a similar magnitude, the majority of which occur along the west coast.
He said: "The movements are a relic of a post-glacial uplift left over from the last ice age.
"The fault line is probably about 100m long and will have moved by about 0.5cm, so it's very minor and will not have even caused superficial damage, although people will have experienced their windows rattling.
"If we compare it to the earthquake in Japan, which had a magnitude of 9, this Blackpool quake is billions of times smaller in terms of energy.
"Only when a quake reaches a magnitude of 4 do we see any superficial damage."
An earthquake measuring 2.4 was recorded near Inverness on Tuesday with several villagers in Abriachan reporting they heard "a rumble".
The largest recorded earthquake experienced in the UK occurred in 1931 and measured 6.1.
The epicentre was Dogger Bank in the North Sea, meaning it had little impact on the mainland.
Seismologists say many of the quakes in the UK are clustered around an enormous block of rock known as the Midlands Microcraton.
This is an ancient, Precambrian (older than 590 million years) feature that runs up through Birmingham towards Stoke-on-Trent.
It is composed of harder rocks than those either side of it and scientists believe it is likely that, in response to tectonic pressures originating in the Atlantic, where the surface of the earth is being pulled apart, those softer rocks on either side are disturbed.
There are a number of active faults that line the Midlands Microcraton and many of the tremors experienced on the western side of England, up to the Pennines, are a result of rocks jostling in this area.


French authorities are seeking €356m (£276m) in unpaid taxes from Booking.com, according to documents filed by parent company Priceline.

Documents filed with US regulator said French authorities recently completed an audit of Booking.com's accounts from 2003 to 2012.
The French government said Booking.com had a base in France and was obliged to pay income and value-added taxes.
The company said the majority of funds being sought are penalties.
"In December 2015, the French tax authorities issued Booking.com assessments for approximately €356m, the majority of which would represent penalties and interest," the filing with the Securities and Exchange Commission said.
The company said believed it complied with local tax law, and would contest the ruling in court if it could not reach a settlement with the French government.
In the same filing Priceline said Italian tax authorities were examining "whether Booking.com should be subject to additional tax obligations in Italy".
Last week, Google's headquarters in Paris were searched as part of an investigation into possible tax evasion.


Roger Bennett grew up in Liverpool in the 1980s with posters of Ferris Bueller, The Fridge and Bob Latchford on his bedroom wall.

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So it is hardly surprising he would find himself in the US in the 1990s, looking for work and trying to find a way to watch Everton games.
"When I first moved here I would go to a bar on Saturday mornings and there would be about a dozen Brits there," said Bennett, who got his dad to hold the radio up to the phone so he could listen to Everton's 1995 FA Cup semi-final win over Spurs (he was presumably too broke to repeat the trick for the final).
"We got one game a week then and it would often be something awful like Sheffield Wednesday versus Leicester.
"Every now and then an American would wander in and ask 'is Full-ham on?' To which the chilling answer from the scariest-looking Brit would be 'walk on, mate, walk on.'
"The impression was that global football was a lecture room and Americans were lucky to be allowed in and they had to keep their hands down.
"Now they have full-throatedly fallen in love with football and it is a joyous thing."
The evidence of that was clear at the Brooklyn Expo Center, where Bennett and fellow ex-pat Michael Davies were about to hold their most ambitious lecture yet: a two-day celebration of what most of the attendees who I spoke to called "soccer…sorry, football".
There was a queue of 1,300 customers around the building, most of them wearing English football club shirts or scarves, bartenders lining up pints of IPA and stout in the beer tent, and staff at the food stall filling the heated shelves with chicken and mushroom pies.
But more startling than this giddy outbreak of Anglophilia in what is meant to be New York's coolest neighbourhood was the crowd gathered in the VIP room.
Because there - in the borough that is home to the NBA's Brooklyn Nets and NHL's New York Islanders and which gave birth to American sporting icons like the Dodgers, Michael Jordan and Vince Lombardi - were the boss of the Premier League and the chairmen or chief executives of four of his member clubs.
Oh, and the bosses of the USA's Major League Soccer and Germany's Bundesliga were on their way.
"This is about how much America loves world football, but also how much world football loves America," said Bennett, the best teacher a new student of the game could ever hope for, as he announced the start of Blazercon, the football convention spawned by his and Davies' phenomenally successful Men in Blazers podcast and TV show.
Equal parts Saint and Greavsie, Wayne's World and Alistair Cooke's Letter from America, Men in Blazers has gone from a conversation at a wedding about how much they miss football banter to becoming a weekly television show on NBC Sports and what Bennett describes as "a beachhead" for the wave of invaders from across the Atlantic.
"America getting football is what clubs in Europe have been dreaming about for decades," Bennett said.
"This is the final frontier and it is a fight for bandwidth."
Much like marriages, partners in double acts know when to jump in with a helpful explanation.
"There have been fans here for years," said Davies, a south Londoner who came to the States 25 years ago to play tennis but became a smash hit at producing TV shows and is now the Ernie Wise of US sports broadcasting's funniest duo.
"But many fans here are just falling in love with it now. That means they don't necessarily think Man Utd and Liverpool are that good.
"They might prefer Southampton or Crystal Palace or Bournemouth. It's a free-for-all!"
That sense of the American football market being a Wild West must be what brought the chairmen of those three clubs to Blazercon, and they were all in the front row when Premier League executive chairman Richard Scudamore opened the convention with a very relaxed question-and-answer session.
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Scudamore has run England's top flight since 1999 but he knows the US well having visited often as a child and then later working for a newspaper group in New York, which explains how he flummoxed the Blazercon staff by arriving at the venue via the subway. They had been looking out for "the commissioner's motorcade".
But there was nothing understated about the numbers he delivered: the Premier League's live TV audiences in the US are up 150% since 2013 and there are now 60 million American adults who have seen at least one game of professional club soccer from somewhere in the world on TV.
"Football has had its ups and downs here but it really does seem to be on the rise and it's very encouraging, very pleasing and very exciting," Scudamore told me before his Q&A.
"Already the audiences here are averaging what you'd get in UK for a Sky or BT game.
"Of course, it's a bigger country. But in terms of unique adults, we had about 32 million Americans watch a game last season, compared with 14-15 million in the UK.
"So our audiences are big and growing. The day more people will be watching live here than in the UK will happen."
Even with the disparity in populations, that will be significant milestone for football fans still regularly lampooned in the UK as Ted Lasso-types who want Boston United to win the "Soccerbowl" and cannot pronounce Leicester City.
But Scudamore is right: it will happen, and soon.
If you include the large numbers of fans who watched the games together, most experts think more than 30 million Americans watched the national team's games against Portugal and Belgium at the 2014 World Cup.
When the US hosted the World Cup in 1994, Bennett watched the games on obscure cable-TV channels with the Latino busboys in bars and restaurants. Nobody else cared.
So what has changed?
"Globalisation is the starting point," said Scudamore.
"People have got less and less parochial in their tastes - they like the best of everything. Sport, generally, has got more global."
He added that it was always a matter of time before the large number of boys and girls who play football in the US translated into an interest in the professional game, a trend that has been greatly helped by the internet.
"[US rights-holder] NBC says about 65% of their output is traditional TV, there is another segment watching on their PCs and there's another segment watching on tablets or mobile," he explained.
"But I think it's more than that. This conference is a testament to what can go on around the main show.
"It's about interaction and social media: it's about people wanting to consume a whole dialogue around the main event. Maybe football is the digital sport."
Scudamore was the warm-up act on Blazercon's first night to former Leyton Orient owner Barry Hearn. The US loves the former Leyton Orient owner and pub-sport impresario.
"I think there are a few reasons why this has happened now," said Hearn.
"There is a problem with head injuries in American football. Maybe parents are looking at that.
"Then there's the Hispanic interest - they have a lot of knowledge about football.
"But most important of all is that football is a soap opera. There are characters, huge money, lifestyle, aspirations, people from nothing who have become millions… there's a story."
As he said this, two Americans in Orient tops introduced themselves. They had decided to become Orient fans after hearing Hearn on a Men in Blazers podcast and had taken advantage of his offer to leave tickets at the Brisbane Road box office for visiting "Great Friends Of the Pod", or GFOPs as they are known on the Blazercon beachhead.
"But the great thing is that these guys support clubs," Hearn continued.
"They decide they want to be Orient fans or Aston Villa fans or Arsenal and so on. They are buying into football in the community.
"I think the new TV deal has helped by levelling the playing field out. So the advantages that Manchester United and Liverpool had, even Newcastle with their huge support, are lessened because of TV revenues.
"Clubs like Southampton and Leicester, with good management, can become major teams. That's quite attractive to the American market, because they love an underdog."
Southampton chairman Ralph Krueger, a Canadian who played ice hockey in Germany and coached in the NHL, agreed with Hearn about the opportunities North America offered clubs like his.
Krueger was almost evangelical when he told me about the plans he is hatching for Southampton FC soccer schools in the US and Canada.
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But then his counterpart at Bournemouth, Jeff Mostyn, was equally excited about the prospect of opening American branches of the Cherries supporters club, and Crystal Palace co-chairman Steve Parish was positively purring at the imminent arrival of American investment at Selhurst Park.
I could not help wondering whether this newfound enthusiasm for the US was a reaction to ignoring it for so long.
"If you go back over the last seven or eight years, I don't think anybody in the Premier League would have predicted that this would become a key market," admitted Liverpool chief executive Ian Ayre, who works for the club's American owners.
"Everybody always knew the opportunity in the US was huge but I think everybody is trying to catch up with the pick-up it's had in the last five or six years."
Was this a result of focusing too much on the exponential but harder-to-pin-down riches on offer in Asia?
"I think it is true about Asia but that's because it's the biggest game there and it was a natural fit. We've got millions of fans across Asia and have been touring there for years," said Ayre.
"But in recent times we've seen the growth here. We came here a couple of years ago and filled Yankee Stadium. We played at Fenway Park and the crowds just get bigger and bigger, the appetite gets bigger and bigger."
Hearn, who has sold snooker to China and darts to Singapore, distilled it even further.
"I have a rule in life: never ignore any market, any customer," he said.
"The North American market is good now but could be great. The Asian market is great but could be even better.
"The secret is to take it all, everywhere - don't leave a penny on the table."
Hearn is very unlikely now to ever own a Premier League team (although he still owns Orient's ground) but it is obvious that his rules in life have some currency around top-flight boardrooms.
For whatever reason - the all-pervasive popularity influence of EA Sport's Fifa games, the fact that Premier League games only have to compete with cartoons on Saturday mornings, not America's traditional big leagues, or what Brad Friedel described to me as the "fast and frenetic" product that appeals to American tastes - football has arrived in the States and it has a surprisingly familiar accent.
"I find it ridiculous," admitted Davies. "But I sort of understand it."


An appeal to find 18 witnesses to provide new evidence for the Hillsborough inquests has seen half identified in the first 24 hours.

The appeal was launched at the request of coroner Sir John Goldring and the victims' families.
Of 6,500 witnesses interviewed so far, investigators had been unable to trace those captured on camera despite "exhausting" all other avenues.
Two police officers, one steward and six fans are still to be traced.
Ninety-six football fans died after the crush at the 1989 FA Cup semi-final between Liverpool and Nottingham Forest at Sheffield Wednesday's Hillsborough stadium.
One witness, who was on the pitch when the disaster unfolded, came forward on Thursday after identifying himself from a photograph.
An Operation Resolve spokesman said eight more had now been identified, including a Nottingham Forest fan.
He said: "The response has been fantastic. To get 50% in the first 24 hours is really good."
Those with information have been asked to contact Operation Resolve on 08000 283 284.
The match between was abandoned at 15:06 BST on 15 April 1989.
Fresh inquests being held in Warrington, Cheshire, continue.


The government has announced it is re-examining its policy on transgender prisoners following the death of a woman in a male prison in Leeds.

Vikki Thompson was being held at Armley where she was found dead on Friday. She told her friends she would kill herself if she was sent to a male prison.
The incident prompted the prisons minister to respond to an urgent question in the House of Commons.
Andrew Selous told a debate they are "reviewing prison service instruction".
Updates on this story and more from around West Yorkshire
Tim Farron, Lib Dem leader, tweeted: "We must reform the law urgently."
Cat Smith, shadow minister for women and equalities, asked if the policy guidelines for transgender prisoners were adequate, and if they are being properly applied.
Mr Selous responded: "Her death like all others in custody is a tragedy and we are totally committed to reducing the number of deaths in prisons.
"We believe that Vikki Thompson was being looked after in accordance with the relevant procedures, but that is now a matter for the Prisons and Probation Ombudsman and Coroner."
He said a review of the policy relating to the care of transgender prisoners had begun earlier this year, and revised policy guidance will be issued and implemented "in due course".
It is estimated there are around 80 transgender prisoners in the UK, although the Ministry of Justice does not keep precise figures.
The debate also follows a transgender woman being sent to a men's prison in Bristol.
80
is the current estimate of transgender people in UK prisons
85,977 prisoners are men, according to the latest figures
3,935 prisoners are female
Tara Hudson, from Bath, was later transferred to a female prison after a campaign to get her moved.
Ms Smith said: "I first raised the issue of Tara Hudson, a young trans woman who was placed in a men's prison on the floor of this house on the 3rd of November.
"It's a tragedy that within three weeks of that, we are once again discussing the issue of trans prisoners."
Jackie Brooklyn, Ms Hudson's mother, said her daughter was "absolutely heartbroken and disgusted" when she heard about Vikki Thompson's death.
She said: "I just think if she was living as a woman she should have been put in a ladies' jail, possibly under segregation for her welfare."
Greg Mulholland, Liberal Democrat MP for Leeds North West, said: "It really seems like something from another era to be putting a transgender woman into a man's prison.
"The tragic consequences of that is there needs to be an inquiry into how this happened. It's certainly time to change the guidelines to avoid this happening again."
Thompson, from Keighley, West Yorkshire, was handed a 12-month jail term in August which was suspended for 24 months.
The BBC understands she later breached the terms of the sentence and was remanded in custody at Bradford Crown Court.
An investigation into her death has been launched, the Prison Service said.
West Yorkshire Police said her death was not being treated as suspicious.
An inquest into Vikki Thompson's death is expected to be opened next week in Wakefield.
Source: The Care and Management of Transsexual Prisoners, Ministry of Justice website


An asteroid that measures nearly 2.7km (1.7 miles) across has flown past the Earth.

The space rock, which is called 1998 QE2, is so large that it is orbited by its own moon.
It made its closest approach to our planet at 20:59 GMT (21:59 BST), but scientists had said there would be no chance it would hit.
Instead it kept a safe distance - at closest, about 5,800,000 km (3,600,000 miles).
That is about 200 times more distant than the asteroid "near-miss" that occurred in February - but Friday's passing space rock is more than 50,000 times larger.
Prof Alan Fitzsimmons, an astronomer at Queen's University Belfast, said: "It's a big one. And there are very few of these objects known - there are probably only about 600 or so of this size or larger in near-Earth space.
"And importantly, if something this size did hit us one day in the future, it is extremely likely it would cause global environmental devastation, so it is important to try and understand these objects."
This fly-by gave astronomers the chance to study the rocky mass in detail.
Using radar telescopes, they were due to record a series of high-resolution images.
They want to find out what it is made of, and exactly where in the Solar System it came from.
Prof Fitzsimmons said: "We already know from the radar measurements, coupled with its brightness, that it appears to be a relatively dark asteroid - that it's come from the outer part  of the asteroid belt."
Early analysis has already revealed that the asteroid has its own moon: it is being orbited by another smaller piece of rock that is about 600m (2000ft) across.
About 15% of asteroids that are large are "binary" systems like this.
This celestial event was not visible to the naked eye, but space enthusiasts with even a modest telescope might be able to witness the pass.
After this, asteroid 1998 QE2 will hurtle back out into deep space; Friday's visit will be its closest approach for at least two centuries.
Researchers are becoming increasingly interested in potential hazards in space.
So far they have counted more than 9,000 near-Earth asteroids, and they spot another 800 new space rocks on average each year.


The family of a man who was killed when a tyre exploded in his face say they have been through 10 years of "hell".

Matthew Hoare, 21, of Chartham, Kent died in January 2006 while working for Watling Tyres in Canterbury.
The company has pleaded guilty at Canterbury Crown Court to health and safety breaches.
His mother Carolyn Hoare said: "I feel like my heart has been ripped out. There's not a moment I don't think about him."
Mr Hoare had been sent to fix a puncture on an earth mover in Sandwich when the tyre blew up.
In April 2013 an inquest recorded a verdict of accidental death.
In January 2015 it was announced Watling Tyres would face three charges of breaching the Health and Safety at Work Act and Working Time Regulations.
Mrs Hoare said: "You live it every day - it doesn't go away. You go to bed thinking about it and you dream during the night.
"I'm living it every day - it's been hell."
Watling Tyres will be sentenced at a later date.


Bristol De Mai romped home to win the Peter Marsh Chase at Haydock as the well-touted Alary disappointed.

The six-year-old victor, ridden by Noel Fehily, was cut to about 16-1 from 66-1 for the Cheltenham Gold Cup in March.
Alary, trained by Colin Tizzard, had been considered a Gold Cup hope but was pulled up before the third last fence.
Bristol De Mai sealed a Haydock double for Nigel Twiston-Davies after The New One became the first horse to win the Champion Hurdle Trial three times.
The nine-year-old, ridden by Sam Twiston-Davies for his trainer father, produced a gutsy display to edge past runner-up Clyne.
Unbeaten Neon Wolf ran out a nine-length victor of the novices' hurdle, while 2014 Champion Hurdle winner Jezki returned from a 632-day absence with a comfortable success at Navan.
Earlier, Ascot's Grade One meeting and Taunton's card on Saturday were called off because of frozen ground.
The Ascot fixture was due to feature the Clarence House Chase, which has been rescheduled to take place at Cheltenham on Festival Trials Day on 28 January.
Cheltenham will now have a nine-race card next week, with racing starting at midday.
Cornelius Lysaght, BBC horse racing correspondent
Abandoned Ascot was billed as the day's top fixture, but had it been on, it'd have struggled to compete for interest with Haydock in terms of the build-up to March's Cheltenham Festival.
The Nigel Twiston-Davies-trained pair The New One and Bristol De Mai showed off the fact they are players - not necessarily the biggest - but definitely players in whatever they go for at Cheltenham.
But the horse to really catch the eye was the Harry Fry-trained Neon Wolf who breezed home under jockey Noel Fehily (2-05) and will line-up in one of the Festival's big novice hurdles.


There are few better places to get away from it all than in the hills of Wales, but which summit is the most remote for solitude seekers?

A new booklet, Y Pellennig: The Remotest Hills of Wales, is attempting to answer that very question.
Topping the list in Wales is West Tump, which is a 17m (55ft) high rock out in the Celtic Sea.
On mainland Wales, the 467m (1,532ft) high Tyle Garw in the Brecon Beacons is 4.9km (3 miles) from the nearest road.
Qualification to be listed among Y Pellennig - which translates from Welsh as far or outlying - requires the hills to be a minimum of 2.5km (1.5 miles) from summit to the nearest road and a minimum of 15m (50ft) of drop from the summit to the lowest point connecting the hill to next higher ground.
The booklet, by Myrddyn Phillips and Aled Williams, provides a hill list that will take the user to some of the most beautiful land in Wales.
Mr Phillips said: "It is a unique concept and challenge as no grouping of hills have been listed by remoteness and no known person has visited all the summits.
"Each hill name has been painstakingly researched by Aled Williams taking in all available Ordnance Survey maps, old estate survey maps, nautical charts and many local enquiries.
"Many of the islands listed are out of bounds during the summer due to seabird nesting colonies, or their general inaccessibility."
The full list and all the details of Y Pellennig are now available online.


The sexual grooming of children has been condemned by Muslim leaders across the UK in a sermon read to thousands of worshippers.

Organisers Together Against Grooming (TAG) said imams at hundreds of mosques had pledged to read the sermon to congregations during Friday prayers.
The sermon highlighted how the Koran emphasised that Muslims must protect children and the vulnerable.
The policing minister Damian Green said it was a "very important" move.
"It reminds people that the vast majority, the overwhelming majority, of British Muslims, condemn child sexual abuse as strongly as any other group in modern Britain," he said.
Sheikh Ibrahim Mogra, an imam at Abu Bakr Mosque in Leicester, said: "People were troubled by us reading the sermon and one man asked me how he could stop it being read.
"He said 'it was not our fault this had happened, our religion does not teach us to do these things and we are condemning it'.
"But as I said to him our only option is to speak out about it."
Mr Mogra added the sermon's message was very clear "this is an evil against humanity" and he was "absolutely delighted with the response".
On Thursday, seven men who abused girls as part of a sadistic sex grooming ring based in Oxford were jailed for life at the Old Bailey. Two of the men were of east African origin and five of Pakistani origin.
By Ed ThomasNorth of England Correspondent, BBC News
In the Lister Hill area of Bradford the Friday call to prayer was the same as usual.
What was different at the Islam Bradford Centre was the sermon, or Khutbah, delivered by the imam.
Aylas Karmani had a message to deliver about the street grooming of young vulnerable girls.
It followed the jailing of men in Oxford and similar cases in Telford, Rochdale and Rotherham.
Many of those involved were British Pakistanis.
Today's sermon talked about Islam being a religion that protects women and children. Those who abused young girls were called evil.
Nobody here said grooming is just a British-Pakistani or Muslim problem.
The majority of child sex offenders are white but what organisers hoped was today would show a united front from Muslim communities around the country.
Hundreds of mosques were asked to read out the sermon.
Many will have done so, but what we don't know is if those who groom will listen.
The Muslim Council of Britain, the Mosque and Imams National Advisory Board and the Islamic Society of Britain all pledged to devote sermons to the issue of sexual grooming, said TAG, a not-for-profit organisation set up to tackle sexual grooming in the UK.
However, Monawar Hussain, founder of The Oxford Foundation, which runs educational programmes to promote religious and social harmony, said the sermon was a "fundamental error of judgement" that would play into the hands of far-right groups.
Mr Hussain, imam at Eton College, said: "Our view is that there is no Muslim on the face of this earth that does not already know that child sexual abuse is evil and wicked: this is normal standard teaching in most mosques.
"There is a terrible danger that far-right groups will point to this and say 'I told you so'."
The sermon, written by Alyas Karmani, an imam and youth worker in Keighley, West Yorkshire, opened with a quotation from the Koran forbidding "sexual indecency, wickedness and oppression of others".
These "disgraceful actions" must be wholeheartedly condemned, it added.
It finished with a call for action and reminds Muslims to speak out if they see any "evil action".
Mr Karmani said: "There's a profound disrespect culture when it comes to treating women. One of the reasons we feel this is the case is poor role models.
"Access to pornography, which also objectifies women, is creating a culture where men are now ambiguous when it comes to the issue of violence against women."
Speaking before the sermon was read out at Friday prayers, Mr Karmani said it had been circulated in an effort to counter what he claimed was a taboo in mosques about talking about sex.
The sermon is the first phase of a "hard-hitting" campaign following a number of high-profile child grooming cases involving Asian men in Bradford, Oxford, Rochdale and Telford, said TAG spokesman Ansar Ali.
"We have been horrified by the details that have emerged from recent court cases and, as Muslims, we feel a natural responsibility to condemn and tackle this crime," said Mr Ali.
We wholeheartedly condemn the disgraceful actions of those  involved in these cases and welcome the convictions in the cases that have been through the courts.
We wish to show our support for the victims of this terrible crime, many of whom are innocent children and we wish to affirm that Islam as a religion of mercy and compassion places a strong obligation on safeguarding and protecting the weak and vulnerable from oppression and abuse particularly of women and children.
Full text of sermon
"Potentially on a Friday you've got hundreds of thousands of people walking into a mosque and you have their undivided attention, so what better medium to try and send a powerful message and raise awareness?"
While sexual grooming and child abuse affected all sections of society and was perpetrated by people of all ethnic groups, the Koran exhorted Muslims to "act against evil and injustice and create just societies", he added.
"We are united in our stand against sexual grooming and, as Muslims, we are leading the effort to rid society of this crime."
The Muslim Council of Britain said that, in conjunction with TAG, it had circulated a Khutbah (Friday sermon) to all affiliated mosques and Islamic centres addressing the issue of grooming.
In a statement it said: "The sermon... raises awareness about what has recently been revealed of the horrific cases of abuse, condemns the behaviour and highlights teachings from the Koran, which obligates the safeguarding and protection of women and children."
Former Labour MP for Keighley Ann Cryer said she was "delighted" by the move, which she said showed the issue was being taken more seriously than in the past.
Ms Cryer said she was approached by mothers worried about grooming in 2002, and was frustrated when police, social services and mosque elders took no action.
"I just hope this message gets beyond the mosque to the non-attenders, because by and large the people who behave like this don't go to the mosque," she said.


It was a case of not being able to see the wood for the cars at one of Northern Ireland's top tourist attractions over the Easter weekend.

The reality of the Dark Hedges did not live up to the fantasy for those visiting the tunnel of trees, made famous by Game of Thrones.
There was no chance of an unspoilt picture with cars and coaches blocking the road and honking their horns.
There are plans to close the road to traffic permanently.
Four objections have been raised to the plans and they are currently being considered, the Department of Infrastructure said.
Jonathan Hobbs, who works for voluntary campaign group NI Greenways, caught the chaos on camera when visiting the County Antrim attraction with his family on Easter Monday.
"You were jostling for space with cars and coaches and I could not let the kids out of my sight," he said. "Tourists travel thousands of miles to come to see this great Game of Thrones location.
"It was a horrible environment and not selling the right image of Northern Ireland."
He also expressed concern about the damage being caused to the trees' roots by the volume of traffic.
A hotel on the Dark Hedges estate, which is just a few minutes' walk from the tourist site, posted on its Facebook page that it has plenty of free car parking.
McComb's Coaches said it had discussed using the car park at a nearby golf club but access to the Dark Hedges would involve crossing a road "where cars go at such terrific speed, it would be an accident waiting to happen".
Caroline McComb, one of the company's directors, said unlike most other coach firms, hers would be in favour of closing the road as long as the infrastructure was put in place and the Dark Hedges remained free of charge.
"People absolutely love the Dark Hedges," she told BBC news NI.
"They just want a picture there and it is becoming more and more difficult to get one.
"There's lots of blame being put on the buses and coaches with regards to damage, but it is just not being maintained.
"Someone needs to take ownership or it will get more run down."
Jamie Laverty, 27, who works near the Dark Hedges and is originally from the nearby village of Stranocum, believes the majority of locals would be in favour of closing the road as the tourist attraction brings in revenue, but there would be some local landowners who would have objections.
"I pass the Dark Hedges on my way to work and even early in the morning, there are cars there," he said.
"It is a beautiful natural phenomenon and we need the tourist infrastructure if it is still going to be here in 30 years."
A Department for Infrastructure spokeswoman said the proposed traffic order would ban most vehicles from the Bregagh Road, encompassing The Dark Hedges, between its junctions with Ballinlea Road and Ballykenver Road.
A package of measures, she added, would aim to "provide an improved experience for local people and visitors alike while helping to reduce the environmental impact on the tree tunnel".
Causeway Coast and Glens Borough Council said it was "working with the landowners and Transport Northern Ireland to create a sustainable solution for the Dark Hedges".
The trees became famous around the world after they featured as the Kings Road in the HBO hit TV series.
There were originally about 150 of the beech trees, which were planted more than 200 years ago by the Stuart family along the entrance to their Gracehill House mansion.
About 90 trees remain, after some were felled dark during recent storms.


Large protests are taking place on the streets on the Philippine capital, Manila, where leaders are attending the Apec regional trade summit.

Hundreds of people from indigenous, student and labour groups clashed with police, who deployed water cannon.
The summit has been overshadowed by territorial disputes over China's activities in the South China Sea.
Leaders have also called for greater global anti-terror co-operation following the Paris attacks.
A draft copy of a declaration due to be released later today says leaders at the Asia Pacific Economic Cooperation (Apec) summit "strongly condemn" all acts of terror and stress the "urgent need for increased international co-operation and solidarity in the fight against terrorism".
Read more: What got Filipinos talking at the Manila meet?
Protesters gathered on Thursday morning in the Manila Bay area, where the Apec meetings have been held.
The anti-globalisation protesters were calling for Apec to be dismantled, accusing the trade bloc of taking advantage of poorer countries.
A protest leader, Renato Reyes, told the Associated Press news agency: "Apec and imperialist globalisation have only benefited the rich countries while further impoverishing developing countries like the Philippines."
Tens of thousands of soldiers and policemen are already in place in the sprawling city to guard against disruptions and potential terrorist attacks.
Demonstrators carrying big placards under the hot sun are chanting anti-Apec, anti-US slogans such as "down with Apec, down with America".
Hundreds of police with batons, truncheons and shields are blocking the protesters from getting near the venue.
Trucks with water cannons are poised to disperse the crowds. Police are currently holding their ground against protesters.
They are also playing loud dance music to drown out the unified voices of the protesters.
World leaders at the summit have been discussing issues such as climate change and regional economic co-operation.
But the dispute among several Asian countries who have overlapping claims in the resource-rich South China Sea has also been in focus.
US President Barack Obama earlier called for China to stop its land reclamation work on disputed reefs. He also signed defence agreements with the Philippines, one of several countries which is in dispute with China on the issue.
Mr Obama said there was a need for "bold steps to lower tensions" in the region.
Chinese President Xi Jinping is also attending the conference but has yet to comment on the issue. China says its work is legal and it has no plans to militarise the new artificial islands.


Examining South Yorkshire Police's handling of the so-called Battle of Orgreave is "a huge, complex logistical exercise", a watchdog has said.

The force referred itself to the IPCC over claims officers involved in the clashes of June 1984, during the UK miners' strike, were told what to write in their statements.
The allegations were revealed in a BBC Inside Out programme in October.
The IPCC has not yet decided whether to launch a full investigation.
On 18 June 1984, British Steel's coking plant at Orgreave was the scene of clashes between about 10,000 striking miners from pits across the country and some 5,000 police officers.
In the Inside Out programme, Vera Baird, Solicitor General during the last Labour government, said police officers were asked by the force's detectives to describe in their statements "scenes they'd simply never seen".
A barrister at the time, Ms Baird first heard the claims during a trial at Sheffield Crown Court in 1985 while she was defending 15 of the 93 pickets arrested at Orgreave.
The trial collapsed after 16 weeks when it became clear police evidence was unreliable.
A spokesperson for the Independent Police Complaints Commission said: "The IPCC has been identifying the location of documentation and preparing plans for conducting a complex scoping exercise to assist it in identifying the specific allegations stemming from events at the Orgreave coking plant and which, if any, warrant investigation.
"We know there are at least 65 boxes of documents, with an estimate of 1,000 documents per box, in an archive in Sheffield.
"This creates a huge, complex logistical exercise, especially for an organisation with limited resources like the IPCC.
"We are committed to concluding this scoping exercise as soon as we can."
An IPCC inquiry is also being carried out into South Yorkshire Police's conduct at Hillsborough.
It was launched after the Independent Hillsborough Panel Report found police and emergency services deflected blame for the disaster on to fans.
The panel revealed 164 police statements had been altered - 116 of them to remove or change negative comments about the policing of the 1989 FA Cup semi-final between Liverpool and Nottingham Forest at the Sheffield stadium, during which a crush on the terraces led to the deaths of 96 Liverpool fans.


Defenders Clint Hill and Philippe Senderos will leave Rangers at the end of the season while midfielder Emerson Hyndman has returned to Bournemouth.

Hill, 38, and 32-year-old Senderos signed one-year deals last summer under Mark Warburton but neither is being retained as Pedro Caixinha looks to rebuild the squad he has inherited.
Hyndman's loan spell has been ended by injury.
He tweeted that he had enjoyed "an incredible time" at the Ibrox club.
The 21-year-old American arrived in January, along with Jon Toral on loan from Arsenal. He made 17 appearances and scored four goals.
Hill was one of 11 signings made by Warburton last summer, as the club prepared for its return to the top-flight.
The Englishman made 31 appearances, scoring six goals, including the equaliser in March's 1-1 draw at Celtic.
Former Arsenal defender Senderos was sent off on his debut in his team's 5-1 Old Firm defeat to Celtic back in September and made three further appearances.


Bordeaux-Begles scored three late tries to send Ulster crashing to defeat in their opening match of the European Champions Cup.

Ulster led for most of the match after Andrew Trimble's 21st-minute try.
But Sebastien Taofifenua battled over to edge the French club ahead eight minutes from the end.
Ulster conceded a penalty try, with Sean Reidy sin-binned for tackling a man without the ball, before winger Blair Connor got the third score.
The pressure will now be on Ulster to beat Exeter in their next Pool 5 fixture on Saturday as another defeat would effectively end their hopes to making the quarter-finals.
They were under considerable pressure in the early stages and trailed to an early Ian Madigan penalty.
But skipper Trimble found a great angle to take Ruan Pienaar's pass and charge through for the opening try midway through the first half.
Jackson converted and added a penalty to give Ulster a 10-3 interval advantage.
Madigan reduced the gap with a second penalty early in the second half and Lionel Beauxis and Jackson traded kicks to leave Ulster 13-9 up.
That is how it stayed until the 73rd minute when a home penalty into the corner set up a five-metre line-out and Taofifenua scored from the driving maul.
Bordeaux were then awarded a penalty try when Reidy's early tackle denied replacement scrum-half Baptiste Serin a score and Beauxis slotted the conversion.
At 21-13 down, Ulster declined a kick at goal when a penalty might have earned them a losing bonus point.
And, to rub salt in their wounds, Brisbane-born winger Connor broke away for the home side's third try seconds before the final whistle.
Ulster's Operations Director Bryn Cunningham: "I am stunned at the moment as at half-time I thought we were in the driving seat. We should have been more out of sight.
"The second half was noting short of atrocious. I am absolutely gobsmacked at how we played to be honest."
Bordeaux-Begles: J Buttin; A Ashley-Cooper, J Dubié, R Lonca, B Connor; I Madigan, Y Lesgourgues; J Poirot, C Maynadier, V Cobilas, J Marais, C Cazeaux, L Jones, H Chalmers, L Goujon.
Replacements: O Avei, S Taofifenua, M Clerc, T Palmer, M Tauleigne, B Serin, L Beauxis, M Talebulamaijaina.
Ulster: J Payne; A Trimble (capt), L Marshall, S Olding, C Gilroy; P Jackson, R Pienaar; A Warwick, R Best, A Ah You, A O'Connor, F van der Merwe, I Henderson, C Ross, S Reidy.
Replacements: R Herring, K McCall, R Kane, P Browne, R Wilson, P Marshall, J Stockdale, R Lyttle.


Leicester City defender Christian Fuchs has signed a new contract to keep him at the club until June 2019.

Fuchs, 30, made 32 appearances last season as the Foxes won the Premier League for the first time.
The former Austria international joined the club on a free transfer from German side Schalke in 2015 and has played 45 times in all competitions.
"Nobody expected what we achieved so far and I think there's still a lot to come," Fuchs told LCFC TV.
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TB Joshua, one of Nigeria's best known preachers, has linked a small plane circling over a church guesthouse in Lagos to its subsequent collapse.

At least 40 people died when the two-storey guesthouse for members of the Synagogue Church of All Nations collapsed on Friday.
There is no official comment. Local officials have said additional floors were being added to the structure.
Mr Joshua - also known as the Prophet - regularly preaches to vast crowds.
The popular televangelist says a small plane had circled over the building four times before it collapsed on Friday afternoon and it may have been an attempt to kill him.
"It was yesterday, Friday, as I was praying; they came to call me that there was an airplane hovering over the church. They said it is like it wanted to land," his statement said.
He said about 30 minutes later, it came over the building several times.
"About five minutes after, the building came down."
He gave journalists a CCTV video clip of the alleged plane, urging them "to use their discretion to write on what they saw".
Rescue officials say 130 people were pulled out of the rubble - some are being treated in local hospitals. Many foreigners are believed to have been there at the time.
Mr Joshua's services, at a mega-church in Lagos's Ikotun district, are known for attracting thousands of people.
Followers in Nigeria and abroad believe he has the gifts of healing and prophecy.


The billionaire owner of Moscow's only fully private airport has been placed under house arrest by a Russian court.

Dmitry Kamenshchik was detained after serving as a witness in an inquiry into a bomb attack at Domodedovo airport in January 2011 which killed 37 people.
Investigators say the airport did not conduct thorough enough passengers searches at the time of the explosion.
He denies the charge, saying regulations then in force did not require such strict controls.
Mr Kamenshchik's fortune has been estimated at $2.9bn (Â£2bn).
In an unexpected twist, government prosecutors disputed the court ruling, calling the accusations unfounded and contesting the plea for house arrest.
Mr Kamenshchik, 47, will remain under house arrest for two months.
The case against him comes against the background of the state's long-running attempts to acquire a stake in Domodedovo, the BBC's Oleg Boldyrev reports from Moscow. Mr Kamenshchik is refusing the sale.


A stray cat hunt is under way in Angus after they were found to be interbreeding with the Scottish wildcat.

A team of trappers has been recruited to catch the feral animals in cages and take them to a vet for neutering.
It is hoped that this will prevent "hybrid" wildcat varieties multiplying.
Dr Roo Campbell, from Scottish Wildcat Action, said mating with domestic cats could diminish the wildcat's ability to survive in the wild.
The pilot scheme in the Angus Glens around the town of Kirriemuir will cost up to Â£20,000 but, if successful, it could be rolled out across parts of the Highlands.
Once found all over Britain, wildcat numbers are dwindling and the species is now restricted to an area north of Scotland's central belt.
They are about 1kg heavier than the domestic cat, with distinctive stripes and large, bushy tail.
Dr Campbell said:"We have known or suspected for years that Scottish wildcats were interbreeding with domestic cats.
"Over time, these hybrid cats living in the wild become less and less wild. They become less well-adapted to their natural environment. It makes them 'soft'."
Dozens of traps will be set within the target area.
The contract guidelines allow for lures like tinned cat food to be used to attract the cat into the cage.


A glasshouse built in honour of British explorer Sir Joseph Banks has been saved from demolition - and will become home to crocodiles and red pandas.

The building was to be pulled down as part of a redevelopment of the Lawn complex in Lincoln.
Bosses at Woodside Wildlife Park have now stepped in to save it and will move it to their site near Langworth.
It will be used as an attraction housing exotic animals and coral reef aquariums.
The conservatory is named after the Lincolnshire botanist who travelled with Captain James Cook on his first voyage to the South Pacific in 1768.
The Grade II listed Lawn complex was sold by City of Lincoln Council last year to the Stokes coffee company which plans to open a cafe and museum on the site.
Neil Mumby, director of the wildlife park, said he wanted to save the building, which housed exotic plants and Koi carp and was popular with generations of families.
"I, like a lot of people, spent my younger days coming here and bringing my children here," he said.
"When I heard that it was being demolished and closing down, I thought we were probably in as good a place as anybody to save it."
Once rebuilt, the conservatory will be used for education and conservation projects, while retaining its heritage as a journey of discovery, Mr Mumby said.
He said the plan to house crocodiles, red pandas and exotic plants inside it would "highlight changes in our planet" since Sir Joseph Banks' voyage of discovery.
The council said it had donated a parcel of land at the Lawn to the Sir Joseph Banks Society to build a new conservatory.


A police force's open letter to a suspected burglar - which included emojis and hashtags - has met with a mixed response online.

Kingston Police tweeted the note addressed to Tracey Dyke, who is suspected of multiple burglaries, accusing her of "blanking" them.
Some users applauded the "novel" social media appeal, while others accused the Met of "public shaming".
Police said Ms Dyke was a suspect in crimes involving "vulnerable victims".
Officers posted the appeal, featuring a photograph of Ms Dyke, and the caption: "Please stop ignoring us Tracey".
In a two-page letter appealing for her to contact detectives, they wrote: "We have come round to see you a number of times recently but it looks like you'd rather not speak to us, which is very disappointing.
"We have a slight suspicion that you might be blanking us #Awkward. You don't text, you don't call back and haven't accepted our friend request."
They added: "We won't stand for this and want to have a discussion with you at our custody suite."
Met Police borough commander Nick Downing congratulated Kingston Police on its "very innovative" appeal.
"Sure the brilliant communities of #Kingston will help track this suspect down," he wrote.
But others were less impressed with the tone of the letter, and accused the force of being "unprofessional" by using hashtags and emoji.
Christina wrote: "This reads like what a school teacher types up on a newsletter to try and be hip and cool but just comes across as awkward".


Dementia patients admitted to hospital in England play "Russian roulette" with their health, a charity is warning.

The Alzheimer's Society said it had found "shocking" evidence of poor and variable care during its review.
The report, based on Freedom of Information (FOI) requests, found problems with falls, night-time discharges and readmissions, and said standards needed to improve urgently.
The Department of Health said the disease was a key priority.
One in four hospital beds is believed to be occupied by a person with dementia.
The Alzheimer's Society called for all hospitals to publish an annual statement of dementia care, to include information on satisfaction, falls, readmissions and staff training.
The charity received responses to their FOI request from half of the 163 hospital trusts in England; however, for some of the questions the figures were based on a fifth of trusts as not all hospitals could provide answers to all the questions.
The information showed:
The Alzheimer's Society also carried out a survey of dementia patients. It found examples of patients being treated with excessive force, not being given enough help with meals and drinks and being left in wet or soiled sheets.
Nine in 10 said hospitals were frightening and only 2% felt all staff understood the needs of people with dementia.
The charity described these findings as unacceptable and a sign that dementia patients were not getting the standard of care they should.
Alzheimer's Society chief executive Jeremy Hughes said: "In the worst cases, hospital care for people with dementia is like Russian roulette. People with dementia and their carers have no way of knowing what's going to happen to them when they are admitted.
"In many cases they are well looked after but, as our investigation shows, too often people with dementia fall and injure themselves, get discharged at night or are marooned in hospital despite their medical treatment having finished."
A Department of Health spokesman said the disease was a key priority and in recent years Â£50m had been spent on making hospitals and care homes more "dementia friendly", while 500,000 staff had received extra training.
"People with dementia and their carers deserve the very best support," he added.


Lewis Hamilton says his Mercedes team are the "underdogs" in their battle with Ferrari heading into this weekend's Bahrain Grand Prix.

The Briton shares the championship lead with Ferrari's Sebastian Vettel after a win and a second place apiece in the first two races of the season.
"Ferrari are going to be very, very strong here. They always have been," Hamilton said.
"It is going to be hard to keep with them, particularly in hot conditions."
Hamilton's concerns stem from tyre usage in the desert heat.
Compared to Ferrari, Mercedes struggled on the softer tyres used in the first race of the season in Australia, when the temperatures went up on race day.
The heat in Bahrain is much more intense than in Melbourne, even though the race starts at twilight and runs into darkness.
Hamilton said: "Ferrari have shown tremendous pace and I think in the first race, particularly in race trim, they're very, very quick - particularly on the hotter circuit.
"So, being that it is a hotter race here, I think the gap is going to be even closer.
"They are always generally the team that is lighter on tyres. We have a heavier car; they have a light, nimble car."
Hamilton said it was "pretty cool" that his former team-mate and long-time rival Fernando Alonso was racing at the Indianapolis 500 this May instead of the Monaco Grand Prix.
But he underlined the challenge facing the two-time champion at the race, which is held at a super-speedway oval.
"It's a completely different category; it's a lot, lot different," Hamilton said.
"I've never driven one of those tracks, but it would be so much different for us to go and do that - the banking, how much you lift, how you use the tyres.
"It takes us guys forever through these test days to learn how to operate these tyres in the right window, so I would imagine it's not an easy thing to do in one go.
"So it's definitely a tall order in a short space of time, but you've got one of the best drivers in the world going there.
"For sure, he will be the best driver in the paddock, but he won't have the experience any of them have. So I think it will be a very exciting challenge for him."
Alonso said: "I need to adapt very quickly. It is very challenging and it is going to be very difficult but that is exciting as well."
The Spaniard has come up with the plan with McLaren and engine partner Honda because he has an ambition to become only the second man in history after Graham Hill to win the so-called triple crown of the Monaco Grand Prix, Indy 500 and Le Mans 24 Hours.
The 35-year-old said winning this year was "not the target", but added: "At the same time, if I want to win the triple crown, one day I have to win it. So let's do it as soon as possible."
Alonso's contract with McLaren runs out this season and he said the team's desire to keep him for 2018 was not part of the discussions over Indianapolis.
"After the summer I will think and I will make a decision of what I will do next year, so nothing has changed after this discussion," he said.
"It's not that I'm thinking 'away from Formula 1'. I'm closer and closer to McLaren and to renew the contract so it's completely independent. After the summer I will see."
He added: "My commitment and my desire to win in F1 is bigger than any other challenge that I can find outside F1 so it's F1 or nothing at the moment."
Alonso is planning to visit the next IndyCar race in Alabama next weekend and have a seat fitting in the Andretti team's factory before flying to the Russian Grand Prix.
He will then spend some time on a simulator before the Spanish Grand Prix, after which he will fly direct to Indianapolis to start practice on the Monday.


Uber has defended itself following accusations that a lack of internal security allowed employees to spy on rides in real-time.

The firm's former forensic investigator made the allegations in a court declaration.
Uber said it had "hundreds of security and privacy experts working around the clock" to protect its data.
The firm recently settled a case in New York that alleged its "God View" tool was used to monitor rides.
"Uber's lack of security regarding its customer data was resulting in Uber employees being able to track high-profile politicians, celebrities, and even personal acquaintances of Uber employees, including ex-boyfriends/girlfriends, and ex-spouses," wrote former forensic investigator Ward Spangenberg in his court declaration.
Mr Spangenberg was fired from Uber earlier this year and is now suing the company over age discrimination and whistleblower retaliation.
Five former security professionals told news site Reveal and the Center for Investigative Reporting that Uber allowed broad access to ride data.
"Uber continues to increase our security investments and many of these efforts, like our multi-factor authentication checks and bug bounty program, have been widely reported," a spokesman for Uber said in a statement.
"This includes enforcing strict policies and technical controls to limit access to user data to authorised employees solely for purposes of their job responsibilities, and all potential violations are quickly and thoroughly investigated."
The spokesman also denied that "all" or "nearly all" Uber's employees had access to customer data, with or without approval.
"Some teams have never had access to this information," he said.
Staff access to data was logged and routinely audited, the spokesman added.
Uber also said that "God View" no longer existed, and it now had an internal tool called "Heaven View".
In January, Uber paid $20,000 (Â£13,700) and promised to strengthen privacy policies in order to settle an investigation by New York's attorney general.
It followed a BuzzFeed news story in 2014 that alleged one of its reporter's rides had been tracked via "God View" without her permission.


Footballer Nile Ranger was seen booking a room at the hotel where, a night earlier, he is accused of raping a woman, a court has been told.

The 22-year-old former Newcastle United striker denies raping a woman he had been out with in Newcastle last January.
The woman said she saw Mr Ranger, who now plays for League One Swindon Town, when she went to see the hotel's CCTV.
Mr Ranger has claimed the woman consented to sex.
She told the court after swapping messages with the footballer, she met him in a bar and next remembered waking up naked in a hotel room the following morning.
During cross-examination by Toby Hedworth QC, defending, the woman, who gave evidence via a videolink, wept when he suggested she had sex twice with Ranger and was "enthusiastically participating".
She has claimed she was too intoxicated to consent and was not attracted to the footballer.
She told the jury she returned to the Carlton Hotel in Jesmond, the following night with a relative to see if they could check the CCTV on the premises.
The alleged victim told the court: "Nile Ranger was at the front desk. He was booking another room for that night, for the same thing to happen to someone else.
"I know this is not the first time this has happened. Since this case, two other people have mentioned being in a horrible state in the same sort of circumstances as what I have been in.
"I'm not going to let it happen to someone else."
Mr Hedworth responded: "You can say what you like about him. You can say he's done it to that person, this person, the other person in the hope that mud sticks."
The alleged victim replied: "That's not true. I know me, I know what happened that night."
Having seen CCTV images of herself looking intoxicated, she said: "Judging by those videos, I'm in no fit state to consent to anything."
Mr Hedworth asked about chatty text messages the pair swapped the morning after the alleged attack, including jokes about Ranger's dance moves, and she replied: "I don't know why I responded like that. I kick myself for responding like that.
"Unless you are in this situation, no-one has a right to say how I dealt with the situation."
The trial continues.


Turkey says it will take a more active role in efforts to end the war in Syria and accepts President Bashar al-Assad as an interim but not long-term player.

But announcing the Turkish policy shift, Prime Minister Binali Yildirim insisted that Mr Assad "can't have a role in Syria's future".
"He is one of the actors today, whether we like it or not," Mr Yildirim said.
Fighting has continued in the city of Hassakeh, where Syrian government jets have bombed Kurdish areas for two days.
Hassakeh, 80km (50 miles) south of the Turkish border, is mainly controlled by the Kurdish YPG militia, but Mr Assad's government forces are trying to push them out.
Turkey is staunchly opposed to Mr Assad, but is also fighting the Kurdistan Workers' Party (PKK) - an ally of the YPG.
Turkey does not want the Kurds to consolidate any territorial gains that could help the PKK.
"In the six months ahead of us, we shall be playing a more active role," Mr Yildirim said. "It means not allowing Syria to be divided along ethnic lines."
A future political settlement for Syria must not include Mr Assad, the PKK or "Daesh" - the so-called Islamic State (IS) group - he said.
Turkey v Syria's Kurds v Islamic State
What is left of Syria after five years of war?
Syrian Kurds declare federal system
Turkey is backing rebel groups fighting to oust Mr Assad, though it has not committed regular troops to the war in Syria. Turkey is hosting about 2.7m Syrian refugees.
Earlier this month Turkey patched up a bitter quarrel with Russia, whose air force is providing vital support for Mr Assad with daily air strikes against the rebels.
The UN Children's Fund (Unicef) on Saturday called for urgent action to help more than 100,000 children "trapped in the horror" of Aleppo, ravaged by intense fighting and heavy air raids by Syrian government and Russian planes.
The children's plight was symbolised this week by the pictures of Omran Daqneesh, an injured five-year-old boy rescued from the rubble in Aleppo. The images of him with a head wound, looking utterly bewildered and shocked, stirred international outrage.
The London-based Syria Solidarity Campaign reported on Saturday that Omran's 10-year-old brother Ali had died of his injuries, after the family home was hit by a bomb. Activists in Aleppo also reported Ali's death, al-Arabiya news said.
The YPG has emerged as a major fighting force in northern Syria in the past two years, becoming a key ally of the US-led coalition against IS.
Kurds made up between 7% and 10% of Syria's population of 24.5 million before the uprising against President Assad began five years ago.


Everton's chief executive has blamed Liverpool City Council for a lack of progress on building a new stadium, saying the club "cannot do it on its own".

Robert Elstone said the club needed the council to commit to investing in the project before it could go ahead.
Plans for a new ground at Walton Hall Park were announced in September 2014.
But Liverpool's mayor said he had yet to receive any detailed proposals for the project.
Responding on Twitter, Joe Anderson wrote: "Re EFC, CEO comments at AGM that they are ready on stadium. I am looking forward to receiving their planning and financial proposals tomorrow."
In a statement he added: "This is a council that helped Everton by purchasing [training ground] Finch Farm. We are trying to find a solution, so to say we are not working in partnership is an insult.
"We are still willing to talk but to say they have not got true partnership is not helpful."
Mr Elstone was speaking to shareholders at Liverpool's Philharmonic Hall, and said without the local authority it would be hard to bring the "challenging funding project" to fruition.
"This is about regeneration of north Liverpool. It's a big prize for Everton but an even bigger prize for the city."
He said the council had failed to recognise the project was a "transformational opportunity".
It was "not an Everton project" and the authority would generate a profit from its lending, he added.
"If the onus is on us to deliver it we can't do it in isolation."
Mr Elstone also told the meeting Everton had reduced its net debt by over Â£17m to Â£28.1m in 2013-14.
Everton's 41,000-capacity Goodison Park is one of the oldest football stadiums and has been the club's home since it opened in 1892.


Premier League Bournemouth survived a scare against Preston to reach the League Cup fourth round on penalties.

Shaun MacDonald's looping header gave Bournemouth the lead but substitute Jordan Hugill nodded the Championship side level to force extra time.
The Cherries took the lead again through Marc Pugh's close-range strike before Daniel Johnson levelled from the penalty spot.
Johnson had the decisive penalty saved by Adam Federici in the shootout.
Hugill and Alan Browne also had penalties saved, with home keeper Jordan Pickford denying Dan Gosling, as Bournemouth won the shootout 3-2.
The Lilywhites, who knocked Premier League Watford out in the previous round, were deservedly behind at the break after MacDonald took advantage of some poor defending to open the scoring.
Simon Grayson's men rallied after the break, though, and Hugill got on the end of Greg Cunningham's cross from the left to head a leveller with six minutes left.
Pugh was at the centre of the action in extra time, stabbing home after Josh King's shot was blocked, before Johnson punished his handball from the penalty spot.
Bournemouth manager Eddie Howe:
"Adam Federici is an outstanding goalkeeper and he's made some great saves there that's got us through and the lads have shown great composure to score at the other end,
"He's an outstanding, experienced international goalkeeper who we certainly believe in.
"He's also a great character and brilliant with the lads, even though he's not playing at the moment he's certainly giving competition for places."
Preston boss Simon Grayson:
"They showed a lot of character and worked extremely hard and showed all of the things that we asked of them after the weekend with that second-half performance.
"I'm pleased with them and disappointed to go out but if you're going to lose a game and respond after a disappointing performance that was the way to do it tonight."
Match ends, Preston North End 2(2), Bournemouth 2(3).
Penalty Shootout ends, Preston North End 2(2), Bournemouth 2(3).
Penalty saved! Daniel Johnson (Preston North End) fails to capitalise on this great opportunity,  left footed shot saved  in the bottom left corner.
Goal!  Preston North End 2(2), Bournemouth 2(3). Marc Pugh (Bournemouth) converts the penalty with a right footed shot to the bottom left corner.
Penalty saved! Alan Browne (Preston North End) fails to capitalise on this great opportunity,  right footed shot saved  in the bottom left corner.
Penalty saved! Dan Gosling (Bournemouth) fails to capitalise on this great opportunity,  right footed shot saved  in the bottom left corner.
Goal!  Preston North End 2(2), Bournemouth 2(2). Kyel Reid (Preston North End) converts the penalty with a left footed shot to the top right corner.
Goal!  Preston North End 2(1), Bournemouth 2(2). Steve Cook (Bournemouth) converts the penalty with a right footed shot to the bottom left corner.
Goal!  Preston North End 2(1), Bournemouth 2(1). Will Keane (Preston North End) converts the penalty with a right footed shot to the bottom left corner.
Goal!  Preston North End 2, Bournemouth 2(1). Junior Stanislas (Bournemouth) converts the penalty with a right footed shot to the centre of the goal.
Penalty saved! Jordan Hugill (Preston North End) fails to capitalise on this great opportunity,  right footed shot saved  in the centre of the goal.
Penalty Shootout begins Preston North End 2, Bournemouth 2.
Second Half Extra Time ends, Preston North End 2, Bournemouth 2.
Corner,  Bournemouth. Conceded by Calum Woods.
Attempt missed. Jordan Hugill (Preston North End) right footed shot from the centre of the box is just a bit too high.
Daniel Johnson (Preston North End) wins a free kick in the defensive half.
Foul by Junior Stanislas (Bournemouth).
Goal!  Preston North End 2, Bournemouth 2. Daniel Johnson (Preston North End) converts the penalty with a right footed shot to the bottom left corner.
Penalty conceded by Adam Smith (Bournemouth) with a hand ball in the penalty area.
Alan Browne (Preston North End) wins a free kick on the left wing.
Foul by Dan Gosling (Bournemouth).
Alan Browne (Preston North End) wins a free kick on the right wing.
Foul by Junior Stanislas (Bournemouth).
Attempt missed. Alan Browne (Preston North End) right footed shot from outside the box is just a bit too high.
Attempt missed. Joshua King (Bournemouth) right footed shot from the left side of the six yard box is close, but misses to the left.
Corner,  Bournemouth. Conceded by Calum Woods.
Foul by Tom Clarke (Preston North End).
Lee Tomlin (Bournemouth) wins a free kick in the defensive half.
Attempt missed. Daniel Johnson (Preston North End) right footed shot from outside the box is just a bit too high.
Foul by Alan Browne (Preston North End).
Lee Tomlin (Bournemouth) wins a free kick in the defensive half.
Second Half Extra Time begins Preston North End 1, Bournemouth 2.
First Half Extra Time ends, Preston North End 1, Bournemouth 2.
Corner,  Bournemouth. Conceded by Jordan Pickford.
Attempt saved. Lee Tomlin (Bournemouth) right footed shot from the centre of the box is saved in the centre of the goal.
Attempt saved. Lee Tomlin (Bournemouth) right footed shot from the centre of the box is saved in the centre of the goal.
Will Keane (Preston North End) is shown the yellow card for a bad foul.
Dan Gosling (Bournemouth) wins a free kick in the defensive half.
Foul by Will Keane (Preston North End).
Corner,  Bournemouth. Conceded by Bailey Wright.


Seven out of 10 under-age drinkers caught in the Cornish holiday resort of Newquay were given alcohol by their parents, according to police.

Most of the teenagers caught drunk and disorderly said they were sent to Cornwall with alcohol, rather than attempting to buy it in local shops.
Supt Julie Whitmarsh from Devon and Cornwall Police described the statistics as "scary".
The force has called for a "national debate" about parental responsibility.
It said although some parents were shocked about their children's behaviour, others  verbally abused officers who confiscated the alcohol.
Many of the 15 and 16-year-olds stopped had been allowed to travel to Cornwall alone with friends to celebrate the end of exams.
More than 4,000 under-18s visited Newquay over the summer, according to figures from a Newquay Safe Partnership campaign to prevent underage drinking.
During Operation Brunel, officers confiscated 1,044 unopened bottles and cans of alcoholic drink and poured away more than 5,000 that were open.
The force said it was talking to the Home Office about ways of prosecuting parents who stock their children up with drink before they leave home.
Supt Whitmarsh said drinking on holiday with peers, without parental control, was a "different situation" than having alcohol at home.
"At what point is it a good idea to leave children with alcohol," she said.
In the most serious cases, parents were telephoned and told to collect their children while a further 71 families were sent warning letters.
Supt Whitmarsh said one officer was verbally abused by a parent who accused him of stopping her son "having fun". The teenager was one of four 16-year-olds caught with 64 cans of Special Brew.
Another mother dismissed an officer's concerns, claiming her child was "not the usual riff-raff".
Police said these were examples of parents not taking their role seriously.
Newquay Safe Partnership was formed in 2009 after the deaths of two teenagers.
Andrew Curwell, 18, from Saddleworth in Greater Manchester, was found at the foot of cliffs at Great Western Beach on 28 June and the following week, 16-year-old Paddy Higgins, from Wokingham in Berkshire, died after falling from cliffs above Tolcarne Beach.


A major festival of light art is to be staged in London next year by the people who organise the biennial Lumiere festival in Durham.

London's first Lumiere festival will feature illuminations and animations projected at 20 locations in the West End and Kings Cross over four nights.
Julian Opie will be among more than two dozen artists taking part.
The Durham version has been running since 2009 and attracted 175,000 people in 2013.
Almost 180,000 turned out when producers Artichoke staged a Lumiere in Londonderry when it was UK City of Culture the same year.
The London event will take place from 14-17 January. Mayor Boris Johnson said it would be "a dazzling new festival for the capital" that would "expel the post-Christmas blues".
The first artists to be confirmed for London are:
Artichoke director Helen Marriage said: "Lumiere was born in Durham and its incredible success has been recognised across the world. Lumiere London will be a very different event with its own artistic programme, but the principles remain the same."
This year's Lumiere will take place in Durham from 12-15 November.


A company that was given a £39m deal to buy timber from Natural Resources Wales - which no other firm was allowed to bid for - has failed to meet a key part of the agreement.

The unnamed saw mill firm had promised to build a new production line for cutting timber.
But the body said the company had not met a deadline to do this by March.
The 10-year deal which was criticised by auditors will now end as a result, it told BBC Wales.
Nick Ramsay, chairman of public accounts committee, said it was "very concerned" by the latest developments.
Auditor General for Wales, Huw Vaughan Thomas, had dubbed the transactions related to the deals for larch and spruce timber "irregular" when he qualified the organisation's accounts in a report that emerged in March.
He said the decision-making process was not transparent and expressed "doubt" over whether it met EU state aid rules.
Around 700,000 cubic meters of timber has been supplied to the firm since 1 April 2014.
As a condition of the contracts, given as part of an effort to deal with diseased larch, the company said it would construct and operate a new saw line by 31 March 2016.
That was missed, and despite the deadline being extended to last Friday, no saw line has been built.
Natural Resources Wales (NRW) told BBC Wales that it was in discussion with the operator about the next steps. It declined to name the contractor.
A spokesman for the Wales Audit Office (WAO) said that one of the "key reasons" NRW entered into the contracts was to create new capacity in the timber industry with a new saw line.
"If sufficient capacity has been achieved without the construction of a new saw line, it does raise further questions about the decision to award the contract to the saw mill operator," the WAO spokesman said.
Llanelli Labour AM Lee Waters, who is a member of the assembly public accounts committee, said the fact NRW has confirmed that the saw mill plan has not happened "draws into question NRW's judgement, and undermines the whole reason for taking this very unusual step in the first place".
Conservative economy spokesman Russell George said the revelation was "extremely damaging" for the organisation.
The assembly's public accounts committee heard from NRW chief executive Emyr Roberts in March, who told AMs that the deal was a "success".
Mr Ramsay said the committee was "not convinced by some of the answers given" by NRW at the previous meeting.
He said the public accounts committee was "very concerned at the latest developments around Natural Resources Wales and its awarding of timber contracts highlighted by the Auditor General for Wales".
"We intend to question NRW closely again next month about its approach to both the awarding of the contracts and of the wider timber market in Wales," he added.
A NRW spokeswoman said: "The prime objective of NRW at the time was to secure a market for the diseased larch."
The spokeswoman denied that NRW's judgement had been drawn into question or that its reasoning for awarding the contracts had been undermined by the failure to open the saw mill line.
"There was insufficient harvesting and processing capacity in Wales at the time and NRW was unable to sell all the infected timber it needed to market," she said.
"The investments identified in the contract were to create the processing capacity for additional private sector larch that was expected to reach the market in addition to NRW's infected timber.
"The company has made investments at the plant and has kept pace with the volumes from NRW managed forests."
She said if larch disease had spread as previously predicted, the additional capacity would have played an important role.
She said the forest industry will be told about how the remaining volume of timber will be offered for sale.
A Welsh Government spokesman said it was disappointed the NRW accounts were qualified and was working with NRW to address the issues raised.


Jordan Spieth says he will "not get ahead" of himself despite holding a three-shot lead going into the final round of The Open at Royal Birkdale.

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The two-time major winner says he will draw on his experience at the 2016 Masters, when he squandered a five-shot lead with nine holes remaining.
Spieth's five-under par 65 on Saturday put him on 11 under, three ahead of American compatriot Matt Kuchar.
"I've had the high and the humbling," said world number three Spieth, 23.
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Victory on Sunday would make Spieth only the second player after Jack Nicklaus, who won a record 18 majors, to have won three of the game's four biggest prizes before the age of 24.
"That would be incredible," he added. "But I've had a five-shot lead in a major and squandered it before. I will keep my head down and not get ahead of myself."
Spieth is the only player to finish every round under par this week and could become the youngest Open winner since a 22-year-old Seve Ballesteros in 1979.
He added: "Everything I've gone through, both good and bad, is advantageous for me. If I win tomorrow, I'll still be learning and it all goes into my process.
"Tomorrow will be emotionally draining but I need to stay neutral in my head."
Iain Carter, BBC Sport golf correspondent at Royal Birkdale
The one thing that needs to be remembered is Spieth had a five-shot lead going into the back nine of the Masters last year.
It was impossible to see that collapse then as it is to see now. We are seeing a remarkable career developing.
Northern Ireland's Rory McIlroy had been three under through five holes, but dropped shots on the seventh and eighth, and a double bogey on the 10th saw him card a disappointing 69.
"Usually you come off Royal Birkdale shooting under par and you would be pretty pleased," said McIlroy, who is tied for 11th on two under.
"But it was so benign out there today and you needed to shoot at least four of five under par to feel like you were making ground out there - and I didn't."
The world number four added: "It's hard to think 'big picture' now, I'm just off the golf course and I'm a little disappointed.
"I definitely feel like today was an opportunity lost to get right in the mix going into tomorrow."
World number 78 Ian Poulter could not hide his frustration following his one-over-par 71, summing up his feelings with a number of expletives as he spent around a minute with reporters.
"I made one bogey yesterday in all those treacherous conditions we had and then I made five bogeys today," said Poulter, 41.
"It's not good enough, simple as that. There were too many stupid mistakes."
The Englishman came through a qualifier at Woburn to win a place at The Open and sat three shots off the pace going into the third round.
But a run of three bogeys on holes 10, 11 and 12 followed a birdie on nine which he hoped would have got "some adrenaline going".
He added: "It's disappointing, extremely disappointing - easy conditions and the greens were soft. No excuses. From a three-putt at the first, which was poor... it's a real shame."
Kuchar, 39, made his Open debut at Royal Birkdale in 1998 and is back bidding for his first top-five finish at The Open at the 12th attempt.
The American was two shots behind playing partner Spieth at the start of the third round, moving within one after a birdie at 14, before a double bogey on 16 helped Spieth move clear.
"I am excited to be in the position I'm in," said Kuchar.
"Three shots off the lead, I've got a great chance with a good day to win the Claret Jug.
"There aren't many other players in the field, even if they have a great final day, who can say that."


A proposal will be put forward to all National League member clubs to change to six-team play-offs at the Annual General Meeting on 10 June.

Currently in non-league's top flight, the four clubs finishing directly below the one automatic promotion place play two-legged semi-finals before a final.
Under the proposals, teams down to seventh in all three of the National League's divisions would be involved.
At the AGM, clubs can vote to amend the proposals, decline them or accept them.
Forest Green won this season's National League promotion final, beating Tranmere Rovers at Wembley 3-1.
FC Halifax Town and Ebbsfleet United won the National League North and South finals respectively, after one-legged semi-final victories.
It is understood that under the proposals all play-off matches will be played over a single leg.
Teams that finish second and third would progress to a semi-final at their home ground, while the teams in fourth and fifth would host seventh and sixth clubs respectively in a "qualifying round".
The winners of those matches would then move into the semi-finals before a final - at Wembley for the National League and a club ground for the North and South divisions.
In theory, that would give teams that finish higher up the division an advantage, as they would play one match less and have a home tie.


A St Andrews University student has told a jury she was terrified when a man pushed her to the ground and whispered "pretty lady" in her ear.

The 19-year-old was walking from halls to a bus stop when she was grabbed.
She was giving evidence at the trial of Pasquale Galianni, 31, who denies sexual assault with intent to rape in North Haugh on 28 November 2013.
Mr Galianni also denies raping a woman and sexually assaulting another woman in St Andrews in April 2014.
He has lodged a special defence of consent to the rape charge.
In evidence  at the High Court in Glasgow, the 19-year-old told advocate depute David Taylor, prosecuting, that it was late afternoon and dark when she was attacked.
She said: "I was walking through a shady area with trees. I noticed someone walking towards me. I was looking at my phone. He seemed to rush towards me, grabbed me and pulled me down to the ground."
The court was told the woman was dragged off the path onto a grassy area. She said that she only got a brief glimpse of her attacker because she was looking at her phone.
The teenager, who cannot be named for legal reasons, said: "I just remember him saying 'pretty lady'. I thought he sounded foreign. It wasn't a British accent. "
When asked how she felt, she said: "Terrified. He was very strong and I didn't think I would be able to push him off."
The student said she tried to make a call to her boyfriend on her mobile phone during the incident and added: "I screamed and basically after I'd screamed he got up and ran off in the direction I'd just come from."
The teenager said she phoned a university friend who came and collected her and took her to the halls of residence where they waited for the police to arrive.
The trial before Lord Burns continues.